Rv2739c Resolved · high auto-curated

H37Rv Rv2739c · MTBC0 mtbc0_002914 · 388 aa · 3074438–3075604 (-) · RefSeq NP_217255.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)transferase
MTBC0 PGAP re-annotationglycosyltransferase
Revised (this work)Glycosyltransferase. Pfam: Glyco_tran_28_C (PF04101.23), EryCIII-like_C (PF06722.19), UDPGT (PF00201.25).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O33282 TrEMBL · unreviewed · Evidence at protein level
UniProt namePossible alanine rich transferase

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category C Energy production and conversion
G Carbohydrate transport and metabolism
eggNOG descriptionglycosyl transferase
Orthologous groupCOG1819

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.82 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 5 synonymous, 10 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Glyco_tran_28_CPF04101.23 8.1e-08285–369 Glycosyltransferase family 28 C-terminal domain
EryCIII-like_CPF06722.19 1.9e-09286–385 Erythromycin biosynthesis protein CIII-like, C-terminal domain
UDPGTPF00201.25 6.1e-06295–363 UDP-glucoronosyl and UDP-glucosyl transferase

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: ephG (epoxide hydrolase), high confidence from genomic context alone (score 789 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv2738c hyp hypothetical protein 884 885 ctx neighborhood:879
Rv2740 ephG epoxide hydrolase 789 789 ctx neighborhood:788
Rv3909 hyp hypothetical protein 750 751 ctx cooccurence:747
Rv3835 hyp hypothetical protein 688 689 ctx cooccurence:683
Rv2843 hyp hypothetical protein 664 665 ctx cooccurence:661
Rv3755c hyp hypothetical protein 643 644 ctx cooccurence:642
Rv0206c mmpL3 transmembrane transport protein MmpL3 640 640 ctx cooccurence:638
Rv2001 hyp hypothetical protein 634 634 ctx cooccurence:634
Rv1111c hyp hypothetical protein 572 572 ctx cooccurence:559
Rv0466 hyp hypothetical protein 545 545 ctx cooccurence:541
Rv2709 transmembrane protein 543 543 ctx cooccurence:543
Rv2736c recX regulatory protein RecX 536 537 ctx neighborhood:532
Rv2737c recA recombinase A 522 522 ctx neighborhood:519
Rv1275 lprC lipoprotein LprC 521 522 ctx cooccurence:520
Rv1752 hyp hypothetical protein 519 520 ctx cooccurence:510

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: transferase
  • MTBC0 PGAP product: glycosyltransferase
  • Pfam (hmmscan --cut_ga): Glyco_tran_28_C PF04101.23 (E=8e-08), EryCIII-like_C PF06722.19 (E=2e-09), UDPGT PF00201.25 (E=6e-06)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217255.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Glyco_tran_28_C (PF04101.23), EryCIII-like_C (PF06722.19), UDPGT (PF00201.25)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1819
  • Curated reference: UniProt O33282 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 56 functional partner(s); context anchor ephG
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_002914|Rv2739c|
MRVAVVAGPDPGHSFPAIALCQRFRAAADTPTLFTGVEWLEAARAAGIDAVELDGLAATDRDLDAGARIHRRAAQMAVLNVPRLRALEPELVVSDVITACGGMAAELLGIPWVELNPHPLYLPSKGLPPIGSGLAAGTGIRGRLRDATMRALTGRSWRAGLRQRAAVRVEIGLPARDPGPLRRLIATLPALEVPRPDWPAEAVVVGPLHFEPTDRVLAIPAGTGPVVVVAPSTALTGTAGLTEVALQSLTPGETVPSGSRLVVSRLSGADLTVPPWAVAGLGSQAELLTRADLVICGGGHGMVAKTLLAGVPMVVVPGGGDQWEIANRVVRQGSAVLIRPLTADALVAAVNEVLSSPRFREAARRAAASVAGAADPVRVCHDALALAG