ephG Resolved · high auto-curated

H37Rv Rv2740 · MTBC0 mtbc0_002915 · 149 aa · 3075648–3076097 (+) · RefSeq NP_217256.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)epoxide hydrolase
MTBC0 PGAP re-annotationepoxide hydrolase
Revised (this work)Epoxide hydrolase. Pfam: LEH (PF07858.19), SnoaL_2 (PF12680.14).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O33283 SwissProt · reviewed · Evidence at protein level
UniProt nameEpoxide hydrolase EphG
EC (curated) EC 3.3.2.10, EC 3.3.2.11
Curated functionEpoxide hydrolase capable of hydrolyzing long or bulky lipophilic epoxides such as 9,10-epoxystearic acid and cholesterol 5,6-oxide in vitro. The physiological substrates have yet to be identified, but could be fatty acid or steroid derivatives.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category Q Secondary metabolites biosynthesis, transport and catabolism
Preferred nameephG
eggNOG descriptionLimonene-1,2-epoxide hydrolase
Orthologous groupCOG4308
EC number EC 3.3.2.8
KEGG orthology K10533
KEGG pathways map00903
Gene Ontology (23) GO:0003674, GO:0003824, GO:0004301, GO:0005575, GO:0005623, GO:0005886, GO:0008150, GO:0008152, GO:0009987, GO:0016020, GO:0016787, GO:0016801 +11 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 1.05 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
LEHPF07858.19 4.1e-6215–135 Limonene-1,2-epoxide hydrolase catalytic domain
SnoaL_2PF12680.14 5.2e-1221–119 SnoaL-like domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv2739c (transferase), high confidence from genomic context alone (score 789 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3618 monooxygenase 926 901 database:900
Rv2739c transferase 789 789 ctx neighborhood:788
Rv2738c hyp hypothetical protein 788 788 ctx neighborhood:787
Rv3169 hyp hypothetical protein 504 505 ctx cooccurence:499
Rv1683 bifunctional long-chain acyl-CoA synthase/lipase 494 494 ctx cooccurence:494
Rv2741 PE_PGRS47 PE-PGRS family protein PE_PGRS47 461 461 ctx neighborhood:461
Rv3805c aftB terminal beta-(1->2)-arabinofuranosyltransferase 422 422 ctx cooccurence:422
Rv0401 transmembrane protein 422 422 ctx cooccurence:422
Rv0913c dioxygenase 416 416 ctx cooccurence:409
Rv1702c hyp hypothetical protein 434 414 ctx cooccurence:412
Rv1148c hyp hypothetical protein 410 411 ctx cooccurence:409
Rv2777c hyp hypothetical protein 410 411 ctx cooccurence:410
Rv3531c hyp hypothetical protein 406 407
Rv3818 hyp hypothetical protein 403 403 ctx cooccurence:403
Rv1945 hyp hypothetical protein 421 401

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: epoxide hydrolase
  • MTBC0 PGAP product: epoxide hydrolase
  • Pfam (hmmscan --cut_ga): LEH PF07858.19 (E=4e-62), SnoaL_2 PF12680.14 (E=5e-12)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217256.1)
  • Domains: Pfam-A via hmmscan --cut_ga — LEH (PF07858.19), SnoaL_2 (PF12680.14)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG4308
  • Curated reference: UniProt O33283 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 31 functional partner(s); context anchor Rv2739c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_002915|Rv2740|ephG
MAELTETSPETPETTEAIRAVEAFLNALQNEDFDTVDAALGDDLVYENVGFSRIRGGRRTATLLRRMQGRVGFEVKIHRIGADGAAVLTERTDALIIGPLRVQFWVCGVFEVDDGRITLWRDYFDVYDMFKGLLRGLVALVVPSLKATL