Rv2729c Family assigned · low auto-curated

H37Rv Rv2729c · MTBC0 mtbc0_002903 · 301 aa · 3063985–3064890 (-) · RefSeq NP_217245.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)integral membrane protein
MTBC0 PGAP re-annotationDMT family transporter
Revised (this work)DMT family transporter.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O33234 TrEMBL · unreviewed · Predicted
UniProt nameProbable conserved integral membrane alanine valine and leucine rich protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category E Amino acid transport and metabolism
G Carbohydrate transport and metabolism
eggNOG descriptionCOG0697 Permeases of the drug metabolite transporter (DMT) superfamily
Orthologous groupCOG0697

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 1.013 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 5 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3434c (transmembrane protein), medium confidence from genomic context alone (score 624 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1326c glgB 1,4-alpha-glucan branching protein 805 782 coexpression:411 database:572
Rv1562c treZ malto-oligosyltrehalose trehalohydrolase 791 781 coexpression:409 database:572
Rv2529 hyp hypothetical protein 714 704 database:516
Rv2731 hyp hypothetical protein 645 631 ctx neighborhood:570
Rv3434c transmembrane protein 624 624 ctx cooccurence:624
Rv2730 hyp hypothetical protein 606 606 ctx neighborhood:606
Rv2727c miaA tRNA delta(2)-isopentenylpyrophosphate transferase 604 588 ctx neighborhood:558
Rv2728c hyp hypothetical protein 570 570 ctx neighborhood:558
Rv3264c manB D-alpha-D-mannose-1-phosphate guanylyltransferase ManB 586 561
Rv1782 eccB5 ESX-5 type VII secretion system protein EccB5 524 524 ctx cooccurence:513
Rv2726c dapF diaminopimelate epimerase 524 504 ctx neighborhood:503
Rv3809c glf UDP-galactopyranose mutase 512 487 coexpression:470
Rv1328 glgP glycogen phosphorylase 511 481
Rv3465 rmlC dTDP-4-dehydrorhamnose 3,5-epimerase 502 472 coexpression:454
Rv2725c hflX GTP-binding protein HflX 472 472 ctx neighborhood:472

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: integral membrane protein
  • MTBC0 PGAP product: DMT family transporter
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217245.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0697
  • Curated reference: UniProt O33234 (TrEMBL, unreviewed; Predicted)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 39 functional partner(s); context anchor Rv3434c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_002903|Rv2729c|
MASVEFATILALGAALLAGIGYVTLQRSARQVTAEEYVGHFTLFHLSLRHALWWLGSLAAVASFTLQAIALTMGSVVLVQSLQATALLFALLIDARLTHHRCTPREWMWAVLLAGAVAVIVMSGNPAAGTTRAPFSTWAVVAVVVVPAVVLCVVGARIASGSLSAVLLAVASSATLAVFTVLTKGVVTELGEGFATLIRTPELYAWILVLPIGLMLQQSSLRVGALTASLPTITVARPVIASVLGITVLDEVLHTGRVALVALVAAVVVVVVATVALARDEVAMMTVSAGELGAAGQLAVR