Rv2041c Family assigned · medium auto-curated

H37Rv Rv2041c · MTBC0 mtbc0_002174 · 439 aa · 2315140–2316459 (-) · RefSeq NP_216557.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)sugar ABC transporter substrate-binding lipoprotein
MTBC0 PGAP re-annotationsugar ABC transporter substrate-binding protein
Revised (this work)Sugar ABC transporter substrate-binding protein. Pfam: SBP_bac_1 (PF01547.31), SBP_bac_8 (PF13416.12).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O53485 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable sugar-binding lipoprotein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category G Carbohydrate transport and metabolism
eggNOG descriptionBacterial extracellular solute-binding protein
Orthologous groupCOG1653
KEGG orthology K02027
KEGG modules M00207

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.881 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 8 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
SBP_bac_1PF01547.31 3.0e-4952–351 Bacterial extracellular solute-binding protein
SBP_bac_8PF13416.12 8.7e-2856–373 Bacterial extracellular solute-binding protein

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv2039c (sugar ABC transporter permease), high confidence from genomic context alone (score 998 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv2039c sugar ABC transporter permease 999 998 ctx neighborhood:881 cooccurence:759 coexpression:872 experimental:412 textmining:777
Rv2038c ugpC sugar ABC transporter ATP-binding protein 998 992 ctx neighborhood:800 cooccurence:570 coexpression:861 experimental:444 textmining:828
Rv2040c sugar ABC transporter permease 995 987 ctx neighborhood:785 cooccurence:755 coexpression:626 experimental:412 textmining:652
Rv1237 sugB sugar ABC transporter permease SugB 959 943 ctx cooccurence:404 coexpression:449 experimental:828
Rv1236 sugA sugar ABC transporter permease SugA 948 930 ctx cooccurence:431 coexpression:430 experimental:788
Rv2317 uspB sugar ABC transporter permease UspB 956 919 ctx cooccurence:741 coexpression:444 experimental:412 textmining:490
Rv2316 uspA sugar ABC transporter permease UspA 937 908 ctx cooccurence:733 coexpression:428 experimental:412
Rv2834c ugpE sn-glycerol-3-phosphate ABC transporter permease UgpE 929 900 ctx cooccurence:693 coexpression:449 experimental:412
Rv2835c ugpA sn-glycerol-3-phosphate ABC transporter permease UgpA 917 900 ctx cooccurence:708 coexpression:432 experimental:412
Rv2832c ugpC sn-glycerol-3-phosphate ABC transporter ATP-binding protein UgpC 955 860 ctx cooccurence:505 coexpression:468 experimental:444 textmining:695
Rv1238 sugC sugar ABC transporter ATP-binding protein SugC 879 833 ctx cooccurence:444 coexpression:466 experimental:444
Rv2037c transmembrane protein 827 827 ctx neighborhood:774
Rv2042c hyp hypothetical protein 804 804 ctx neighborhood:684
Rv2043c pncA pyrazinamidase/nicotinamidase PncA 693 694 ctx neighborhood:684
Rv1795 eccD5 ESX-5 type VII secretion system protein EccD 627 627 ctx cooccurence:625

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: sugar ABC transporter substrate-binding lipoprotein
  • MTBC0 PGAP product: sugar ABC transporter substrate-binding protein
  • Pfam (hmmscan --cut_ga): SBP_bac_1 PF01547.31 (E=3e-49), SBP_bac_8 PF13416.12 (E=9e-28)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216557.1)
  • Domains: Pfam-A via hmmscan --cut_ga — SBP_bac_1 (PF01547.31), SBP_bac_8 (PF13416.12)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1653
  • Curated reference: UniProt O53485 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 20 functional partner(s); context anchor Rv2039c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_002174|Rv2041c|
MVNKPFERRSLLRGAGALTAASLAPWAAGCAADDDDALTFFFAANPDELRPRMRVVNEFQRRYPDIKVRALLSGPGVMQQLATFCAGGKCPDVLMAWELTYAELADRGVLLDLNTLLARDQAFAAELKSDSIGALYETFTFNGGQYAFPEQWSGNFLFYNKQLFDDAGVPPPPGSWERPWSFAEFLDAAQALTKQGRSGRDRQWGFVNAWVSFYAAGLFAMNNGVPWSVPRMNPTHLNFDHDGFLEAVQFYADLTNKHKVAPSAAEQQSMSTADLFSVGKAGIALAGHWRYQTFDRADGLDFDVAPLPIGPRGRAACSDIGVTGLAIAATSRRKDQAWEFVKFATGPVGQALIGESRLFVPVLRSAINSHGFANAHRRVGNLAVLSEGPAYSEGLPVTPAWEKIAALMDRYFGPVLRGSRPATSLTGLSQAVDEVLRNP