bioA Resolved · high auto-curated

H37Rv Rv1568 · MTBC0 mtbc0_001676 · 437 aa · 1787356–1788669 (+) · RefSeq NP_216084.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)adenosylmethionine--8-amino-7-oxononanoate aminotransferase BioA
MTBC0 PGAP re-annotationadenosylmethionine--8-amino-7-oxononanoate transaminase
Revised (this work)Adenosylmethionine--8-amino-7-oxononanoate transaminase. Pfam: Aminotran_3 (PF00202.28).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WQ81 SwissProt · reviewed · Evidence at protein level
UniProt nameAdenosylmethionine-8-amino-7-oxononanoate aminotransferase
EC (curated) EC 2.6.1.62
Curated functionCatalyzes the reversible transfer of the alpha-amino group from S-adenosyl-L-methionine (SAM) to 7-keto-8-aminopelargonic acid (KAPA) to form 7,8-diaminopelargonic acid (DAPA). It is the only aminotransferase known to utilize SAM as an amino donor. Can also use sinefungin but not S-adenosylhomocysteine as substrate.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category H Coenzyme transport and metabolism
Preferred namebioA
eggNOG descriptionCatalyzes the transfer of the alpha-amino group from S- adenosyl-L-methionine (SAM) to 7-keto-8-aminopelargonic acid (KAPA) to form 7,8-diaminopelargonic acid (DAPA). It is the only animotransferase known to utilize SAM as an amino donor
Orthologous groupCOG0161
EC number EC 2.6.1.62
KEGG orthology K00833
KEGG pathways map00780, map01100
KEGG modules M00123, M00573, M00577
Gene Ontology (47) GO:0003674, GO:0003824, GO:0004015, GO:0006082, GO:0006732, GO:0006766, GO:0006767, GO:0006768, GO:0006790, GO:0006807, GO:0008150, GO:0008152 +35 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.157 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 9 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Aminotran_3PF00202.28 2.4e-11035–428 Aminotransferase class-III

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: bioD (ATP-dependent dethiobiotin synthetase BioD), high confidence from genomic context alone (score 999 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1570 bioD ATP-dependent dethiobiotin synthetase BioD 999 999 ctx neighborhood:881 cooccurence:762 coexpression:695 database:900 textmining:965
Rv1569 bioF1 8-amino-7-oxononanoate synthase 999 998 ctx neighborhood:881 cooccurence:667 coexpression:473 database:900 textmining:946
Rv1589 bioB biotin synthetase 999 976 ctx fusion:633 cooccurence:756 coexpression:693 textmining:963
Rv0032 bioF2 8-amino-7-oxononanoate synthase 989 974 ctx cooccurence:510 coexpression:451 database:900 textmining:596
Rv1571 hyp hypothetical protein 975 882 ctx neighborhood:881 textmining:803
Rv1567c membrane protein 718 719 ctx neighborhood:719
Rv1563c treY maltooligosyl trehalose synthase 635 625 ctx neighborhood:623
Rv1565c acyltransferase 439 439 ctx neighborhood:439
Rv1564c treX maltooligosyl trehalose synthase 440 428 ctx neighborhood:426
Rv1590 hyp hypothetical protein 401 370
Rv2794c pptT 4'-phosphopantetheinyl transferase 668 260 textmining:571
Rv2438c nadE glutamine-dependent NAD(+) synthetase 887 189 textmining:867
Rv0089 methyltransferase 828 134 textmining:811
Rv3602c panC pantothenate synthetase 808 106 textmining:795
Rv3279c birA bifunctional biotin operon repressor/biotin--[acetyl-CoA-carboxylase 768 102 textmining:753

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: adenosylmethionine--8-amino-7-oxononanoate aminotransferase BioA
  • MTBC0 PGAP product: adenosylmethionine--8-amino-7-oxononanoate transaminase
  • Pfam (hmmscan --cut_ga): Aminotran_3 PF00202.28 (E=2e-110)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216084.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Aminotran_3 (PF00202.28)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0161
  • Curated reference: UniProt P9WQ81 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 22 functional partner(s); context anchor bioD
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001676|Rv1568|bioA
MAAATGGLTPEQIIAVDGAHLWHPYSSIGREAVSPVVAVAAHGAWLTLIRDGQPIEVLDAMSSWWTAIHGHGHPALDQALTTQLRVMNHVMFGGLTHEPAARLAKLLVDITPAGLDTVFFSDSGSVSVEVAAKMALQYWRGRGLPGKRRLMTWRGGYHGDTFLAMSICDPHGGMHSLWTDVLAAQVFAPQVPRDYDPAYSAAFEAQLAQHAGELAAVVVEPVVQGAGGMRFHDPRYLHDLRDICRRYEVLLIFDEIATGFGRTGALFAADHAGVSPDIMCVGKALTGGYLSLAATLCTADVAHTISAGAAGALMHGPTFMANPLACAVSVASVELLLGQDWRTRITELAAGLTAGLDTARALPAVTDVRVCGAIGVIECDRPVDLAVATPAALDRGVWLRPFRNLVYAMPPYICTPAEITQITSAMVEVARLVGSLP