Rv1565c Family assigned · medium auto-curated

H37Rv Rv1565c · MTBC0 mtbc0_001673 · 729 aa · 1783604–1785793 (-) · RefSeq NP_216081.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)acyltransferase
MTBC0 PGAP re-annotationacyltransferase family protein
Revised (this work)Acyltransferase family protein. Pfam: Acyl_transf_3 (PF01757.29), SGNH (PF19040.7).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O06625 TrEMBL · unreviewed · Predicted
UniProt nameConserved hypothetical membrane protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category I Lipid transport and metabolism
eggNOG descriptionAcyltransferase family
Orthologous groupCOG1835

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.403 · purifying
Polymorphic sites (≥ 0.1% of strains) 8 synonymous, 9 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Acyl_transf_3PF01757.29 1.4e-2133–377 Acyltransferase domain
SGNHPF19040.7 3.5e-22500–716 SGNH domain (fused to AT3 domains)

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: treX (maltooligosyl trehalose synthase), high confidence from genomic context alone (score 837 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1564c treX maltooligosyl trehalose synthase 843 837 ctx neighborhood:827
Rv1563c treY maltooligosyl trehalose synthase 830 830 ctx neighborhood:827
Rv1562c treZ malto-oligosyltrehalose trehalohydrolase 829 829 ctx neighborhood:827
Rv1566c ripD hyp hypothetical protein 785 778 ctx neighborhood:774
Rv1489A hyp hypothetical protein 527 528 ctx cooccurence:525
Rv0472c HTH-type transcriptional regulator 491 492 ctx cooccurence:476
Rv1354c hyp hypothetical protein 453 452 coexpression:421
Rv3668c protease 452 452 ctx cooccurence:442
Rv1275 lprC lipoprotein LprC 449 449 ctx cooccurence:446
Rv1754c hyp hypothetical protein 447 447 ctx cooccurence:444
Rv1568 bioA adenosylmethionine--8-amino-7-oxononanoate aminotransferase BioA 439 439 ctx neighborhood:439
Rv1510 hyp hypothetical protein 463 429
Rv1814 erg3 membrane-bound C-5 sterol desaturase 419 420 coexpression:401
Rv3630 integral membrane protein 446 415
Rv2972c hyp hypothetical protein 446 411 coexpression:410

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: acyltransferase
  • MTBC0 PGAP product: acyltransferase family protein
  • Pfam (hmmscan --cut_ga): Acyl_transf_3 PF01757.29 (E=1e-21), SGNH PF19040.7 (E=4e-22)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216081.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Acyl_transf_3 (PF01757.29), SGNH (PF19040.7)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1835
  • Curated reference: UniProt O06625 (TrEMBL, unreviewed; Predicted)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 21 functional partner(s); context anchor treX
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001673|Rv1565c|
MLTLSPPRPPALTPEPALPPVTMGTRTTGFYRHDLDGLRGVAIALVAVFHVWFGRVSGGVDVFLALSGFFFGGKILRAALNPDLSLSPIAEVIRLIRRLLPALVVVLAGCALLTIAIQPQTRWEAFANQSLASLGYYQNWELASTVSNYLRAGEAVSPLQHIWSMSVQGQFYLAFLLLVAGCAYLLRRLFRGPRAPYLRTMFVVLLSTLTLASFIYAIVAHHAYQATAYYNTFARAWELLAGALVGAVVPHVRWPMWLRTAVATAALAAILSCGALIDGVKEFPGPWALVPVGATMLMILAGANRQGHPGTRDRLPLPNRLLATAPLVALGAMAYSWYLWHWPLLIFWLSYTGHRHANFVEGAAVLLVSGLLAYLTTRLVEDPLRYRAPAGVRSPAAVPPIPWRLRLRRPTIVLGSVVALLGVALTATSFTWREHVIVQRAAGKELSGLSSRDYPGARALIDHVRVPKLRMRPTVLEVRHDLPTSTKDGCISDFVNPAIINCTYGDVDAPRTIALAGGSHAEHWLTALDLLGRMHHFKVVTYLKMGCPLSTEEVPLIMGNNAPYPQCHQWVQAAMAKLVADHPDYVFTTSTRPWNIKPGDVMPATYVGIWQTFADNNIPVLAMRDTPWLVKDGQPFIPADCLAKGGNPQSCGIARSKVLVDRNPTLDFVARFPLLKPLDMSDAICRTDTCRAVEGNVLVYRDSHHLTPTYMRTMTSELGRQIAANTDWW