Rv2714 Family assigned · medium auto-curated

H37Rv Rv2714 · MTBC0 mtbc0_002888 · 324 aa · 3049480–3050454 (+) · RefSeq NP_217230.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationPAC2 family protein
Revised (this work)PAC2 family protein. Pfam: PAC2 (PF09754.16).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6YA29 TrEMBL · unreviewed · Evidence at protein level
UniProt nameConserved alanine and leucine rich protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionPAC2 family
Orthologous groupCOG2047

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.676 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
PAC2PF09754.16 4.1e-6343–261 PAC2 family

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv2715 (hydrolase), high confidence from genomic context alone (score 803 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv2715 hydrolase 803 803 ctx neighborhood:802
Rv2716 hyp hypothetical protein 754 754 ctx neighborhood:753
Rv2713 sthA pyridine nucleotide transhydrogenase 592 593 ctx neighborhood:591
Rv3260c whiB2 transcriptional regulator WhiB2 563 564 ctx cooccurence:562
Rv3219 whiB1 transcriptional regulator WhiB1 534 534 ctx cooccurence:532
Rv3129 Rv3129, (MTCY164.40), len: 110 aa. Conserved hypothetical protein, with some similarity to various hypothetical proteins from Streptomyces c 530 530 coexpression:450
Rv1321 nucS endonuclease NucS 524 524 ctx cooccurence:521
Rv0004 hyp hypothetical protein 483 483 ctx cooccurence:483
Rv2901c hyp hypothetical protein 477 478 ctx cooccurence:476
Rv2712c hyp hypothetical protein 457 457 ctx neighborhood:457
Rv0080 hyp hypothetical protein 447 447 coexpression:447
Rv1875 hyp hypothetical protein 447 447 coexpression:447
Rv0121c hyp hypothetical protein 447 447 coexpression:447
Rv1331 clpS ATP-dependent Clp protease adapter protein ClpS 428 429 ctx cooccurence:427
Rv1390 rpoZ DNA-directed RNA polymerase subunit omega 428 429 ctx cooccurence:415

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: PAC2 family protein
  • Pfam (hmmscan --cut_ga): PAC2 PF09754.16 (E=4e-63)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217230.1)
  • Domains: Pfam-A via hmmscan --cut_ga — PAC2 (PF09754.16)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2047
  • Curated reference: UniProt I6YA29 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 23 functional partner(s); context anchor Rv2715
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_002888|Rv2714|
MARDQGADEAREYEPGQPGMYELEFPAPQLSSSDGRGPVLVHALEGFSDAGHAIRLAAAHLKAALDTELVASFAIDELLDYRSRRPLMTFKTDHFTHSDDPELSLYALRDSIGTPFLLLAGLEPDLKWERFITAVRLLAERLGVRQTIGLGTVPMAVPHTRPITMTAHSNNRELISDFQPWISEIQVPGSASNLLEYRMAQHGHEVVGFTVHVPHYLTQTDYPAAAQALLEQVAKTGSLQLPLAALAEAAAEVQAKIDEQVQASAEVAQVVAALERQYDAFIDAQENRSLLTRDEDLPSGDELGAEFERFLAQQAEKKSDDDPT