mpg Resolved · high auto-curated

H37Rv Rv1688 · MTBC0 mtbc0_001796 · 203 aa · 1924994–1925605 (+) · RefSeq NP_216204.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)3-methyladenine DNA glycosylase
MTBC0 PGAP re-annotationDNA-3-methyladenine glycosylase
Revised (this work)DNA-3-methyladenine glycosylase. Pfam: Pur_DNA_glyco (PF02245.22).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WJP7 SwissProt · reviewed · Evidence at protein level
UniProt namePutative 3-methyladenine DNA glycosylase
EC (curated) EC 3.2.2.-

UniProt still lists this protein as Putative 3-methyladenine DNA glycosylase; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category L Replication, recombination and repair
Preferred namempg
eggNOG descriptionBelongs to the DNA glycosylase MPG family
Orthologous groupCOG2094
EC number EC 3.2.2.21
KEGG orthology K03652
KEGG pathways map03410
Gene Ontology (31) GO:0003674, GO:0003824, GO:0003905, GO:0006139, GO:0006259, GO:0006281, GO:0006284, GO:0006725, GO:0006807, GO:0006950, GO:0006974, GO:0008150 +19 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 1.12 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Pur_DNA_glycoPF02245.22 1.4e-547–184 Methylpurine-DNA glycosylase (MPG)

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: tyrS (tyrosine--tRNA ligase), high confidence from genomic context alone (score 872 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1689 tyrS tyrosine--tRNA ligase 884 872 ctx neighborhood:872
Rv1687c ABC transporter ATP-binding protein 788 789 ctx neighborhood:787
Rv1686c ABC transporter permease 728 728 ctx neighborhood:726
Rv1685c hyp hypothetical protein 724 724 ctx neighborhood:722
Rv0427c xthA exodeoxyribonuclease III protein XthA 779 673 database:661
Rv1650 pheT phenylalanine--tRNA ligase subunit beta 566 566 ctx neighborhood:544
Rv3859c gltB glutamate synthase large subunit 550 529 ctx neighborhood:529
Rv1317c alkA bifunctional regulatory protein/DNA repair enzyme AlkA 535 172 textmining:462
Rv3586 disA DNA integrity scanning protein DisA 568 156 textmining:510
Rv3674c nth endonuclease III 479 104 textmining:443
Rv0950c hyp hypothetical protein 804 50 textmining:803
Rv2891 hyp hypothetical protein 658 47 textmining:656
Rv3895c eccB2 ESX-2 secretion system protein EccB 658 47 textmining:656
Rv3894c eccC2 ESX-2 type VII secretion system protein EccC 801 44 textmining:801
Rv3786c hyp hypothetical protein 657 44 textmining:656

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: 3-methyladenine DNA glycosylase
  • MTBC0 PGAP product: DNA-3-methyladenine glycosylase
  • Pfam (hmmscan --cut_ga): Pur_DNA_glyco PF02245.22 (E=1e-54)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216204.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Pur_DNA_glyco (PF02245.22)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2094
  • Curated reference: UniProt P9WJP7 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 20 functional partner(s); context anchor tyrS
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001796|Rv1688|mpg
MNAEELAIDPVAAAHRLLGATIAGRGVRAMVVEVEAYGGVPDGPWPDAAAHSYRGRNGRNDVMFGPPGRLYTYRSHGIHVCANVACGPDGTAAAVLLRAAAIEDGAELATSRRGQTVRAVALARGPGNLCAALGITMADNGIDLFDPSSPVRLRLNDTHRARSGPRVGVSQAADRPWRLWLTGRPEVSAYRRSSRAPARGASD