recN Resolved · high auto-curated

H37Rv Rv1696 · MTBC0 mtbc0_001804 · 587 aa · 1931698–1933461 (+) · RefSeq NP_216212.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)DNA repair protein RecN
MTBC0 PGAP re-annotationDNA repair protein RecN
Revised (this work)DNA repair protein RecN. Pfam: SMC_N (PF02463.26).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WHI7 SwissProt · reviewed · Evidence at protein level
UniProt nameDNA repair protein RecN
Curated functionMay be involved in recombinational repair of damaged DNA.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category L Replication, recombination and repair
Preferred namerecN
eggNOG descriptionMay be involved in recombinational repair of damaged DNA
Orthologous groupCOG0497
KEGG orthology K03631
Gene Ontology (6) GO:0005575, GO:0005618, GO:0005623, GO:0030312, GO:0044464, GO:0071944

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.249 · purifying
Polymorphic sites (≥ 0.1% of strains) 6 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
SMC_NPF02463.26 4.3e-152–535 RecF/RecN/SMC N terminal domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: tlyA (16S/23S rRNA (cytidine-2'-O)-methyltransferase TlyA), high confidence from genomic context alone (score 895 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1694 tlyA 16S/23S rRNA (cytidine-2'-O)-methyltransferase TlyA 917 895 ctx neighborhood:867
Rv1695 ppnK inorganic polyphosphate/ATP-NAD kinase 978 882 ctx neighborhood:867 textmining:821
Rv1693 hyp hypothetical protein 981 869 ctx neighborhood:867 textmining:861
Rv1691 hyp hypothetical protein 981 869 ctx neighborhood:867 textmining:866
Rv1692 phosphatase 980 867 ctx neighborhood:867 textmining:861
Rv1698 mctB copper transporter MctB 761 761 ctx neighborhood:756
Rv2737c recA recombinase A 937 708 coexpression:689 textmining:793
Rv1690 lprJ lipoprotein LprJ 668 668 ctx neighborhood:668
Rv1697 steA hyp hypothetical protein 654 654 ctx neighborhood:651
Rv2720 lexA repressor LexA 869 651 coexpression:649 textmining:640
Rv1713 engA GTPase Der 579 579
Rv1650 pheT phenylalanine--tRNA ligase subunit beta 691 569 ctx neighborhood:440
Rv3423c alr alanine racemase 591 551 ctx cooccurence:546
Rv1700 NUDIX hydrolase 549 521 ctx neighborhood:514
Rv1699 pyrG CTP synthase 569 518 ctx neighborhood:513

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: DNA repair protein RecN
  • MTBC0 PGAP product: DNA repair protein RecN
  • Pfam (hmmscan --cut_ga): SMC_N PF02463.26 (E=4e-15)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216212.1)
  • Domains: Pfam-A via hmmscan --cut_ga — SMC_N (PF02463.26)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0497
  • Curated reference: UniProt P9WHI7 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 88 functional partner(s); context anchor tlyA
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001804|Rv1696|recN
MLTELRIESLGAISVATAEFDRGFTVLTGETGTGKTMVVTGLHLLGGARADATRVRSGADRAVVEGRFTTTDLDDATVAGLQAVLDSSGAERDEDGSVIALRSISRDGPSRAYLGGRGVPAKSLSGFTNELLTLHGQNDQLRLMRPDEQRGALDRFAAAGEAVQRYRKLRDAWLTARRDLVDRRNRARELAQEADRLKFALNEIDTVDPQPGEDVALVADIARLSELDTLREAATTARATLCGTPDADAFDRGAVDSLGRARAALQSSDDAALRGLAEQVGEALTVVVDAVAELGAYLDELPADASALDAKLARQAQLRTLTRKYAADIDGVLRWADEARARLAQLDVSEEGLAALERRTGELAHELGQAAVDLSTIRRKAAKRLAKEVSAELSALAMADAEFTIGVTTELADHGDPVALALASGELARAGADGVDAVEFGFVAHRGMTVLPLAKSASGGELSRVMLSLEVVLATSRKQAAGTTMVFDEIDAGVGGWAAVQIGRRLARLARTHQVIVVTHLPQVAAYADVHLMVQRTGRDGASGVRRLTSEDRVAELARMLAGLGDSDSGRAHARELLETAQNDELT