Rv0713 Still unknown · low auto-curated

H37Rv Rv0713 · MTBC0 mtbc0_000755 · 313 aa · 814139–815080 (+) · RefSeq NP_215227.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)transmembrane protein
MTBC0 PGAP re-annotationDUF4436 domain-containing protein
Revised (this work)Conserved hypothetical protein; DUF domain(s) DUF4436. Function unknown.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6WZ58 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable conserved transmembrane protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionDomain of unknown function (DUF4436)
Orthologous group2E1SP

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 1.035 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 9 missense, 0 nonsense, 2 frameshift
Disruption 2 distinct premature-stop/frameshift site(s); most common in 0.47% of strains (683) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
DUF4436PF14494.12 4.3e-10558–311 Domain of unknown function (DUF4436)

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3843c (transmembrane protein), high confidence from genomic context alone (score 723 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3903c cpnT hyp hypothetical protein 744 745 ctx cooccurence:744
Rv3843c transmembrane protein 723 723 ctx cooccurence:723
Rv3166c hyp hypothetical protein 677 677 ctx cooccurence:677
Rv1682 hyp hypothetical protein 669 669 ctx cooccurence:669
Rv3446c hyp hypothetical protein 615 615 ctx cooccurence:613
Rv0538 membrane protein 602 602 ctx cooccurence:602
Rv2164c hyp hypothetical protein 588 588 ctx cooccurence:587
Rv0295c stf0 hyp hypothetical protein 549 550 ctx cooccurence:543
Rv2028c universal stress protein 544 545 ctx cooccurence:543
Rv1775 hyp hypothetical protein 544 545 ctx cooccurence:543
Rv2113 integral membrane protein 515 516 ctx cooccurence:504
Rv3788 hyp hypothetical protein 487 487 ctx cooccurence:474
Rv0712 hyp hypothetical protein 480 481 ctx neighborhood:481
Rv2487c PE_PGRS42 PE-PGRS family protein PE_PGRS42 478 478 ctx cooccurence:478
Rv2423 hyp hypothetical protein 472 473 ctx cooccurence:471

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: transmembrane protein
  • MTBC0 PGAP product: DUF4436 domain-containing protein
  • Pfam (hmmscan --cut_ga): DUF4436 PF14494.12 (E=4e-105)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215227.1)
  • Domains: Pfam-A via hmmscan --cut_ga — DUF4436 (PF14494.12)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 2E1SP
  • Curated reference: UniProt I6WZ58 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 37 functional partner(s); context anchor Rv3843c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000755|Rv0713|
MAGSDPPTGGPASQAGSDAGASPEHKHMSRRKHLVLDVCIILGVLIAYVFSLLGYDWLAHTPGPLPQPDVGTTDDTVVLIRFEELHTVANRLDVKVLVLPDDSMIDHRLQVLTTDTSVRLYPENELGDLQYPVGKLPAQVATTIEAHGNPGAWPFDTYTTDTVQADVLVGAGDNRQYVPARVEVTGSLEGWDISAVRVGESSQTSDRPDNVIITLKRAKGPLVFDLGICLVLITLPTLALFVAIQMITGRRKFQPPFGTWYAAMLFAVVPLRTILPGSPPAGAWIDRAVVIWVLIALAAAMVVYIVAWYRESD