Rv3779 Family assigned · low auto-curated

H37Rv Rv3779 · MTBC0 mtbc0_004007 · 666 aa · 4249106–4251106 (+) · RefSeq NP_218296.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)transmembrane protein
MTBC0 PGAP re-annotationDUF6541 family protein
Revised (this work)DUF6541 family protein.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P72045 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable conserved transmembrane protein alanine and leucine rich

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionPsort location CytoplasmicMembrane, score
Orthologous groupCOG5617

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.513 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 7 synonymous, 10 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
DUF6541PF20176.4 3.0e-8012–656 Family of unknown function (DUF6541)

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: rfbE (O-antigen/lipopolysaccharide ABC transporter ATP-binding protein RfbE), high confidence from genomic context alone (score 882 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3781 rfbE O-antigen/lipopolysaccharide ABC transporter ATP-binding protein RfbE 930 882 ctx neighborhood:881 textmining:434
Rv3780 bpa hyp hypothetical protein 897 882 ctx neighborhood:881
Rv3783 rfbD O-antigen/lipopolysaccharide ABC transporter permease RfbD 930 881 ctx neighborhood:881 textmining:434
Rv3782 glfT1 galactofuranosyl transferase GlfT 908 881 ctx neighborhood:881
Rv3632 membrane protein 893 787 ctx cooccurence:768 textmining:519
Rv0048c membrane protein 773 773 ctx cooccurence:767
Rv0875c hyp hypothetical protein 745 745 ctx cooccurence:743
Rv0517 acyltransferase 737 738 ctx cooccurence:734
Rv3630 integral membrane protein 734 735 ctx cooccurence:718
Rv1510 hyp hypothetical protein 704 705 ctx cooccurence:686
Rv3773c hyp hypothetical protein 703 703 ctx cooccurence:703
Rv1182 papA3 acyltransferase papA3 670 670 ctx cooccurence:670
Rv3446c hyp hypothetical protein 667 667 ctx cooccurence:667
Rv2079 hyp hypothetical protein 661 661 ctx cooccurence:660
Rv3824c papA1 acyltransferase 662 653 ctx cooccurence:653

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: transmembrane protein
  • MTBC0 PGAP product: DUF6541 family protein
  • Pfam (hmmscan --cut_ga): DUF6541 PF20176.4 (E=3e-80)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218296.1)
  • Domains: Pfam-A via hmmscan --cut_ga — DUF6541 (PF20176.4)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG5617
  • Curated reference: UniProt P72045 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 88 functional partner(s); context anchor rfbE
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_004007|Rv3779|
MGLWFGTLIALILLIAPGAMVARIAQLRWPVAIAVGPALTYGVVALAIIPYGALGIPWNGWTALAALAVTCAVATGLQLLLARFRDLDAEALAVSRWPAVTVAAGVLLGALLIGWAAYRGIPHWQSIPSTWDAVWHANTVRFILDTGQASSTHMGELRNVETHAPLYYPSVFHGLVAVFCQLTGAAPTTGYTLSSLAASVWLFPVSAAVLTWRAVRSHPGALWSASCASAEWRAAGAAGTAAALSASFTAVPYVEFDTAAMPNLAAYGIAVPTMVLITSTLRHRDRIPVAVLALVGVFSLHITGGIVVALLVSAWWLFEALRHPVRSRLADLLTLAGVAAMAGLVMLPQFLSVRQQEDIIAGHAFPTYLSKKRGLFDAVFQHSRHLNDFPVQYALIVLAAIGGLILLVKKIWWPLAVWLLLIVMNVDAGTPLGGPIGGVAGALGEFFYHDPRRIAAATTLLLMLMAGVALFATVMLLVAAAKRLTDRFRPQPVSVWASATATLLIGATLVSAWHYFPRHRFLFGDKYDSVMIDQKDLDAMAYLASLPGARDTLIGNANTDGTAWMYAVAGLHPLWTHYDYPLQQGPGYHRFIFWAYGRNGESDPRVLEAIQVLRIRYILTSTPTVRGFAVPDGLVSLETSRSWAKIYDNGEARIYEWRGTAAATHS