rplQ Resolved · high auto-curated

H37Rv Rv3456c · MTBC0 mtbc0_003674 · 180 aa · 3902774–3903316 (-) · RefSeq NP_217973.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)50S ribosomal protein L17
MTBC0 PGAP re-annotation50S ribosomal protein L17
Revised (this work)50S ribosomal protein L17. Pfam: Ribosomal_L17 (PF01196.25).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WHD3 SwissProt · reviewed · Evidence at protein level
UniProt nameLarge ribosomal subunit protein bL17

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category J Translation, ribosomal structure and biogenesis
Preferred namerplQ
eggNOG descriptionribosomal protein L17
Orthologous groupCOG0203
KEGG orthology K02879
KEGG pathways map03010
KEGG modules M00178
Gene Ontology (30) GO:0003674, GO:0003735, GO:0005198, GO:0005575, GO:0005618, GO:0005622, GO:0005623, GO:0005737, GO:0005829, GO:0005840, GO:0005886, GO:0015934 +18 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.189 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 7 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Ribosomal_L17PF01196.25 3.8e-4021–116 Ribosomal protein L17

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: rplO (50S ribosomal protein L15), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0723 rplO 50S ribosomal protein L15 999 1000 ctx cooccurence:608 coexpression:970 experimental:999 textmining:625
Rv0706 rplV 50S ribosomal protein L22 999 1000 coexpression:968 experimental:999 textmining:717
Rv3459c rpsK 30S ribosomal protein S11 999 1000 ctx neighborhood:726 coexpression:936 experimental:999 textmining:622
Rv0700 rpsJ 30S ribosomal protein S10 999 1000 coexpression:968 experimental:999 textmining:434
Rv0979A rpmF 50S ribosomal protein L32 999 1000 coexpression:851 experimental:999 textmining:570
Rv3442c rpsI 30S ribosomal protein S9 999 1000 coexpression:866 experimental:999 textmining:843
Rv0717 rpsN1 30S ribosomal protein S14 999 1000 coexpression:738 experimental:999
Rv0682 rpsL 30S ribosomal protein S12 999 1000 coexpression:946 experimental:999
Rv1643 rplT 50S ribosomal protein L20 999 1000 coexpression:865 experimental:999
Rv1015c rplY 50S ribosomal protein L25/general stress protein Ctc 999 1000 coexpression:834 experimental:999
Rv0634B rpmG2 50S ribosomal protein L33 999 1000 coexpression:730 experimental:999
Rv0720 rplR 50S ribosomal protein L18 999 1000 coexpression:886 experimental:999 textmining:566
Rv2441c rpmA 50S ribosomal protein L27 999 1000 ctx cooccurence:456 coexpression:858 experimental:999
Rv1298 rpmE 50S ribosomal protein L31 999 1000 coexpression:563 experimental:999
Rv3443c rplM 50S ribosomal protein L13 999 1000 ctx cooccurence:606 coexpression:866 experimental:999

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: 50S ribosomal protein L17
  • MTBC0 PGAP product: 50S ribosomal protein L17
  • Pfam (hmmscan --cut_ga): Ribosomal_L17 PF01196.25 (E=4e-40)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217973.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Ribosomal_L17 (PF01196.25)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0203
  • Curated reference: UniProt P9WHD3 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 286 functional partner(s); context anchor rplO
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003674|Rv3456c|rplQ
MPKPTKGPRLGGSSSHQKAILANLATSLFEHGRITTTEPKARALRPYAEKLITHAKKGALHNRREVLKKLRDKDVVHTLFAEIGPFFADRDGGYTRIIKIEARKGDNAPMAVIELVREKTVTSEANRARRVAAAQAKAKKAAAMPTEESEAKPAEEGDVVGASEPDAKAPEEPPAEAPEN