Rv1520 Resolved · high auto-curated

H37Rv Rv1520 · MTBC0 mtbc0_001627 · 346 aa · 1720837–1721877 (+) · RefSeq NP_216036.3

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)sugar transferase
MTBC0 PGAP re-annotationglycosyltransferase
Revised (this work)Glycosyltransferase. Pfam: Glycos_transf_2 (PF00535.33).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WLV5 SwissProt · reviewed · Evidence at protein level
UniProt nameUncharacterized protein Rv1520

UniProt still lists this protein as Uncharacterized protein Rv1520; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category M Cell wall / membrane / envelope biogenesis
eggNOG descriptionGlycosyltransferase like family 2
Orthologous groupCOG0463
Gene Ontology (12) GO:0003674, GO:0003824, GO:0005575, GO:0005618, GO:0005623, GO:0005886, GO:0016020, GO:0016740, GO:0016757, GO:0030312, GO:0044464, GO:0071944

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.378 · purifying
Polymorphic sites (≥ 0.1% of strains) 7 synonymous, 8 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Glycos_transf_2PF00535.33 1.6e-252–132 Glycosyl transferase family 2

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv2957 (PGL/p-HBAD biosynthesis glycosyltransferase), high confidence from genomic context alone (score 761 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1518 hyp hypothetical protein 853 854 ctx neighborhood:449 cooccurence:744
Rv1519 hyp hypothetical protein 787 779 ctx neighborhood:699
Rv2957 PGL/p-HBAD biosynthesis glycosyltransferase 760 761 ctx cooccurence:747
Rv1521 fadD25 fatty-acid--CoA ligase FadD25 490 489 ctx neighborhood:430
Rv1514c glycosyltransferase 475 475
Rv1517 hyp hypothetical protein 447 448 ctx neighborhood:446
Rv0334 rmlA glucose-1-phosphate thymidylyltransferase 460 432
Rv3464 rmlB dTDP-glucose 4,6-dehydratase 438 413
Rv3781 rfbE O-antigen/lipopolysaccharide ABC transporter ATP-binding protein RfbE 421 390
Rv3402c hyp hypothetical protein 401 378
Rv3465 rmlC dTDP-4-dehydrorhamnose 3,5-epimerase 402 371
Rv3824c papA1 acyltransferase 529 80 textmining:510
Rv1182 papA3 acyltransferase papA3 400 80
Rv0254c cobU bifunctional cobinamide kinase/cobinamide phosphate guanylyltransferase 605 55 textmining:600
Rv0247c succinate dehydrogenase iron-sulfur subunit 544 44 textmining:543

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: sugar transferase
  • MTBC0 PGAP product: glycosyltransferase
  • Pfam (hmmscan --cut_ga): Glycos_transf_2 PF00535.33 (E=2e-25)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216036.3)
  • Domains: Pfam-A via hmmscan --cut_ga — Glycos_transf_2 (PF00535.33)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0463
  • Curated reference: UniProt P9WLV5 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 17 functional partner(s); context anchor Rv2957
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001627|Rv1520|
MSIVSISYNQEEYIREALDGFAAQRTEFPVEVIIADDASTDATPRIIGEYAARYPQLFRPILRQTNIGVHANFKDVLSAARGEYLALCEGDDYWTDPLKLSKQVKYLDRHPETTVCFHPVRVIYEDGAKDSEFPPLSWRRDLSVDALLARNFIQTNSVVYRRQPSYDDIPANVMPIDWYLHVRHAVGGEIAMLPETMAVYRRHAHGIWHSAYTDRRKFWETRGHGMAATLEAMLDLVHGHREREAIVGEVSAWVLREIGKTPGRQGRALLLKSIADHPRMTMLSLQHRWAQTPWRRFKRRLSTELSSLAALAYATRRRALEGRDGGYRETTSPPTGRGRNVRGSHA