Rv3402c Family assigned · medium auto-curated

H37Rv Rv3402c · MTBC0 mtbc0_001626 · 50 aa · 1720659–1720811 (+) · RefSeq NP_217919.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationDegT/DnrJ/EryC1/StrS family aminotransferase
Revised (this work)DegT/DnrJ/EryC1/StrS family aminotransferase. Pfam: DegT_DnrJ_EryC1 (PF01041.24).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WGJ7 SwissProt · reviewed · Evidence at transcript level
UniProt nameProtein Rv3402c

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category E Amino acid transport and metabolism
Preferred namevioA
eggNOG descriptionUDP-4-amino-4-deoxy-L-arabinose aminotransferase
Orthologous groupCOG0399
EC number EC 2.6.1.33
KEGG orthology K13308, K20429
KEGG pathways map00523, map01130
KEGG modules M00797

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains) pseudogene candidate

pN/pS 0.292 · purifying
Polymorphic sites (≥ 0.1% of strains) 7 synonymous, 5 missense, 1 nonsense, 0 frameshift
Disruption 1 distinct premature-stop/frameshift site(s); most common in 11.98% of strains (17403) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
DegT_DnrJ_EryC1PF01041.24 4.1e-10150–405 DegT/DnrJ/EryC1/StrS aminotransferase family

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv1503c (Rv1503c, (MTCY277.25c), len: 182 aa. Conserved hypothetical protein, similar to C-terminal region of P27833|RFFA_ECOLI lipopolysaccharide bi), high confidence from genomic context alone (score 717 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv2386c mbtI salicylate synthase 913 831 coexpression:831 textmining:510
Rv2383c mbtB phenyloxazoline synthase 853 823 coexpression:788
Rv1347c mbtK lysine N-acetyltransferase MbtK 863 816 coexpression:810
Rv1349 irtB iron ABC transporter ATP-binding protein/permease IrtB 848 808 coexpression:802
Rv1348 irtA iron ABC transporter ATP-binding protein/permease IrtA 831 787 coexpression:780
Rv2385 mbtJ acetyl hydrolase 785 785 coexpression:785
Rv2384 mbtA 2,3-dihydroxybenzoate-AMP ligase 869 779 coexpression:778 textmining:436
Rv2380c mbtE peptide synthetase 777 777 coexpression:735
Rv2379c mbtF peptide synthetase 764 765 coexpression:732
Rv1505c hyp hypothetical protein 756 741 ctx cooccurence:570
Rv1503c Rv1503c, (MTCY277.25c), len: 182 aa. Conserved hypothetical protein, similar to C-terminal region of P27833|RFFA_ECOLI lipopolysaccharide bi 716 717 ctx cooccurence:715
Rv3404c hyp hypothetical protein 676 657
Rv3784 dTDP-glucose 4,6-dehydratase 605 581 coexpression:478
Rv1500 pimF glycosyltransferase 554 537
Rv1752 hyp hypothetical protein 573 533 coexpression:506

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: DegT/DnrJ/EryC1/StrS family aminotransferase
  • Pfam (hmmscan --cut_ga): DegT_DnrJ_EryC1 PF01041.24 (E=4e-101)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217919.1)
  • Domains: Pfam-A via hmmscan --cut_ga — DegT_DnrJ_EryC1 (PF01041.24)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0399
  • Curated reference: UniProt P9WGJ7 (SwissProt, reviewed; Evidence at transcript level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 33 functional partner(s); context anchor Rv1503c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001626|Rv3402c|
MNAELVESADLTVTEDICSRIVSLPVHDHMAIADVARVVAPFGEGLARGG