Rv0552 Resolved · high auto-curated

H37Rv Rv0552 · MTBC0 mtbc0_000581 · 534 aa · 646440–648044 (+) · RefSeq NP_215066.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationamidohydrolase
Revised (this work)Amidohydrolase. Pfam: Amidohydro_3 (PF07969.18), Amidohydro_1 (PF01979.27).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O06418 TrEMBL · unreviewed · Evidence at protein level
UniProt nameConserved protein

UniProt still lists this protein as Conserved protein; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionamidohydrolase
Orthologous groupCOG1574

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.268 · purifying
Polymorphic sites (≥ 0.1% of strains) 8 synonymous, 6 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Amidohydro_3PF07969.18 3.1e-8350–532 Amidohydrolase family
Amidohydro_1PF01979.27 5.6e-09208–531 Amidohydrolase family

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: menC (muconate cycloisomerase), high confidence from genomic context alone (score 902 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv0553 menC muconate cycloisomerase 902 902 ctx neighborhood:881
Rv0554 bpoC non-heme bromoperoxidase BpoC 878 878 ctx neighborhood:878
Rv0551c fadD8 fatty-acid--CoA ligase FadD8 792 793 ctx neighborhood:784
Rv0557 mgtA GDP-mannose-dependent alpha-mannosyltransferase 761 749 ctx neighborhood:749
Rv0555 menD bifunctional 2-succinyl-6-hydroxy-2,4-cyclohexadiene-1-carboxylate synthase/2-oxoglutarate decarboxylase 683 683 ctx neighborhood:682
Rv0556 transmembrane protein 669 669 ctx neighborhood:669
Rv0548c menB 1,4-dihydroxy-2-naphthoyl-CoA synthase 614 615 ctx neighborhood:615
Rv0074 hyp hypothetical protein 572 572 ctx cooccurence:570
Rv0558 menH demethylmenaquinone methyltransferase 492 492 ctx neighborhood:492
Rv0547c oxidoreductase 405 405 ctx neighborhood:405
Rv2672 protease 688 174 textmining:638
Rv3829c dehydrogenase 401 152
Rv1898 hyp hypothetical protein 808 67 textmining:803
Rv0842 integral membrane protein 715 47 textmining:714
Rv0874c hyp hypothetical protein 806 45 textmining:806

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: amidohydrolase
  • Pfam (hmmscan --cut_ga): Amidohydro_3 PF07969.18 (E=3e-83), Amidohydro_1 PF01979.27 (E=6e-09)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215066.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Amidohydro_3 (PF07969.18), Amidohydro_1 (PF01979.27)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1574
  • Curated reference: UniProt O06418 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 16 functional partner(s); context anchor menC
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000581|Rv0552|
MADADLVMTGTVLTVDDARPTAEAIAVADGRVIAVGDRSEVAGLVGANTRVIDLGAGCVMPGFVEAHGHPLLEAVVLSDRFVDIRPVTMRDADDVVAAIRGEVARRGPAGAYLVGWDPLLQSGLGEPTLTWLDSLAPNGPLVIIHNSGHKAYFNSHAAWLNGLTRDTADPKGAKYGRDGNGELDGTAEEIGAILPLLAGVADPSNFGAMLRAECARLNRAGLTTCSEMAFDPGYRPMVEAVRAELTVRLCTYEISNARMCTDATPGQGDDMLRQVGIKIWVDGSPWVGNIDLTFPYLDTPATRAIGVPPGSRGCANYTREQLAEIVGAYFPRGWQIACHVHGDGGVDTILDVYEEALRRNPRDDHRLRLEHVGAIRPDQLRRAAELGVTCSIFVDQIHYWGDVIVDDLFGAQRGSRWMPAGSAVAAGMRISLHNDPPVTPEEPLRNISVAATRVAPSGRVLAPEERLTVEQAIRAQTIDAAWQLFAEDAIGSLQVGKYADMVVLSADPRTVPPEQIADLAVRATFLAGRQVYRR