Rv0877 Still unknown · low auto-curated

H37Rv Rv0877 · MTBC0 mtbc0_000932 · 262 aa · 979287–980075 (+) · RefSeq NP_215392.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationDUF3027 domain-containing protein
Revised (this work)Conserved hypothetical protein; DUF domain(s) DUF3027. Function unknown. Foldseek best (non-significant) hit: 5kuy-assembly1_G Influenza hemagglutinin H3 A/Hong Kong/1/1968 in comp (prob 0.66, TM 0.50).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WKR3 SwissProt · reviewed · Evidence at protein level
UniProt nameUncharacterized protein Rv0877

UniProt still lists this protein as Uncharacterized protein Rv0877; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionProtein of unknown function (DUF3027)
Orthologous group28NWB

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 1.021 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
DUF3027PF11228.14 1.6e-8245–234 Protein of unknown function (DUF3027)

Structural neighbours (Foldseek on the ESMFold model, exploratory)

ESMFold model confidence: mean pLDDT 91.5 (very high). A confident model makes the fold comparison meaningful.

Best matches against the PDB, ranked by Foldseek homology probability. A high probability / TM-score suggests a shared fold; unless flagged sig (E < 0.01) these are fold hypotheses, not assignments.

TargetProbTME-valueDescription
5kuy-assembly1_G 0.66 0.50 2.4e-01 5kuy-assembly1_G Influenza hemagglutinin H3 A/Hong Kong/1/1968 in complex with designed inhibitor protein HSB.2A
4n6v-assembly7_4 0.44 0.56 1.2e+00 4n6v-assembly7_4 Partial rotational order disorder structure of human stefin B
6i1m-assembly1_A 0.20 0.48 7.4e-01 6i1m-assembly1_A Secreted type 1 cystatin from Fasciola hepatica
6d63-assembly6_L 0.16 0.43 1.2e+00 6d63-assembly6_L The structure of AtzH: a little known member of the atrazine breakdown pathway
6d63-assembly6_K 0.15 0.46 1.9e+00 6d63-assembly6_K The structure of AtzH: a little known member of the atrazine breakdown pathway
6bju-assembly1_A 0.14 0.40 1.3e+00 6bju-assembly1_A The structure of AtzH: a little known member of the atrazine breakdown pathway
6bju-assembly2_C 0.13 0.38 1.3e+00 6bju-assembly2_C The structure of AtzH: a little known member of the atrazine breakdown pathway
1stf-assembly1_I 0.12 0.51 5.4e+00 1stf-assembly1_I THE REFINED 2.4 ANGSTROMS X-RAY CRYSTAL STRUCTURE OF RECOMBINANT HUMAN STEFIN B IN COMPLEX WITH THE CYSTEINE PROTEINASE PAPAIN: A NOVEL TYPE OF PROTEINASE INHIBITOR INTERACTION

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0876c (transmembrane protein), high confidence from genomic context alone (score 776 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv0875c hyp hypothetical protein 884 884 ctx neighborhood:774 cooccurence:506
Rv0876c transmembrane protein 776 776 ctx neighborhood:774
Rv3662c hyp hypothetical protein 635 636 ctx cooccurence:631
Rv3202c adnA ATP-dependent DNA helicase 609 609 ctx cooccurence:608
Rv2699c hyp hypothetical protein 603 604 ctx cooccurence:598
Rv2239c hyp hypothetical protein 599 600 ctx cooccurence:596
Rv0048c membrane protein 517 517 ctx cooccurence:517
Rv1171 hyp hypothetical protein 506 507 ctx cooccurence:501
Rv3721c dnaZX DNA polymerase III subunit gamma/tau 497 498 ctx cooccurence:493
Rv0822c hyp hypothetical protein 483 484 ctx cooccurence:482
Rv1423 whiA transcriptional regulator WhiA 475 476 ctx cooccurence:471
Rv3847 hyp hypothetical protein 455 455 ctx cooccurence:449
Rv2050 rbpA RNA polymerase-binding protein RbpA 453 453 ctx cooccurence:445
Rv2891 hyp hypothetical protein 453 453 ctx cooccurence:451
Rv3597c lsr2 iron-regulated H-NS-like protein 443 444 ctx cooccurence:441

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: DUF3027 domain-containing protein
  • Pfam (hmmscan --cut_ga): DUF3027 PF11228.14 (E=2e-82)
  • Foldseek best: 5kuy-assembly1_G Influenza hemagglutinin H3 A/Hong Kong/1/1968 in complex with d (prob 0.66, E=2e-01, TM=0.50)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215392.1)
  • Domains: Pfam-A via hmmscan --cut_ga — DUF3027 (PF11228.14)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 28NWB
  • Curated reference: UniProt P9WKR3 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Model confidence: ESMFold per-residue pLDDT (mean 91.5, very high)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 23 functional partner(s); context anchor Rv0876c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000932|Rv0877|
MTGPTEESAVATVADWPEGLAAVLRGAADQARAAVVEFSGPEAVGDYLGVSYEDGNAATHRFIAHLPGYQGWQWAVVVASYSGADHATISEVVLVPGPTALLAPDWVPWEQRVRPGDLSPGDLLAPAKDDPRLVPGYTASGDAQVDETAAEIGLGRRWVMSAWGRAQSAQRWHDGDYGPGSAMARSTKRVCRDCGFFLPLAGSLGAMFGVCGNELSADGHVVDRQYGCGAHSDTTAPAGGSTPIYEPYDDGVLDIIEKPAES