Rv3662c Resolved · medium auto-curated

H37Rv Rv3662c · MTBC0 mtbc0_003880 · 256 aa · 4125095–4125865 (-) · RefSeq NP_218179.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationoxidoreductase
Revised (this work)Oxidoreductase.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6YCP7 TrEMBL · unreviewed · Evidence at protein level
UniProt nameFido domain-containing protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionFilamentation induced by cAMP protein fic
Orthologous groupCOG3177

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.799 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 4 synonymous, 8 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: dppD (dipeptide ABC transporter ATP-binding protein DppD), high confidence from genomic context alone (score 883 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv3663c dppD dipeptide ABC transporter ATP-binding protein DppD 928 883 ctx neighborhood:881 textmining:412
Rv3664c dppC dipeptide ABC transporter permease DppC 917 882 ctx neighborhood:881
Rv3665c dppB dipeptide ABC transporter permease DppB 886 808 ctx neighborhood:807 textmining:433
Rv3666c dppA dipeptide ABC transporter substrate-binding lipoprotein DppA 864 808 ctx neighborhood:807
Rv0487 hyp hypothetical protein 743 743 ctx cooccurence:741
Rv3202c adnA ATP-dependent DNA helicase 723 724 ctx cooccurence:702
Rv0877 hyp hypothetical protein 635 636 ctx cooccurence:631
Rv3256c hyp hypothetical protein 623 623 ctx cooccurence:606
Rv1417 membrane protein 586 586 ctx cooccurence:579
Rv3679 anion transporter ATPase 585 585 ctx cooccurence:582
Rv0822c hyp hypothetical protein 568 569 ctx cooccurence:566
Rv2170 GCN5-like N-acetyltransferase 568 569 ctx cooccurence:563
Rv2709 transmembrane protein 559 560 ctx cooccurence:557
Rv1171 hyp hypothetical protein 550 550 ctx cooccurence:537
Rv1638A hyp hypothetical protein 550 550 ctx cooccurence:545

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: oxidoreductase
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218179.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG3177
  • Curated reference: UniProt I6YCP7 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 52 functional partner(s); context anchor dppD
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003880|Rv3662c|
MTVDPLAPLMELPGVAAASDRVRDALSRVHRHRANLRGWPVAAAEASLRAARASSVLDGGPARLHDAGAPTSGKPALSDPVFAGALRVGQALEGGAGPVVGVWRRAPLQALARLHMLAAADQVDDDRLGRPRSDADVGPRLELLADVVTHPTLASAPVVAAVAHGELLTLRPFGCADGVVARAVSRLVTIATGLDPHGLGVPEVIWMRQPAEYHDAARRFAGGTPDGVAGWLLLCCGAMLDGAREALSIAESLSPG