aldA Resolved · high auto-curated

H37Rv Rv0768 · MTBC0 mtbc0_000817 · 489 aa · 865336–866805 (+) · RefSeq NP_215282.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)aldehyde dehydrogenase AldA
MTBC0 PGAP re-annotationaldehyde dehydrogenase
Revised (this work)Aldehyde dehydrogenase. Pfam: Aldedh (PF00171.28).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6X9R9 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable aldehyde dehydrogenase NAD dependent AldA

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category C Energy production and conversion
Preferred namealdA
eggNOG descriptionBelongs to the aldehyde dehydrogenase family
Orthologous groupCOG1012
EC number EC 1.2.1.3
KEGG orthology K00128
KEGG pathways map00010, map00053, map00071, map00280, map00310, map00330, map00340, map00380, map00410, map00561, map00620, map00625, map00903, map00981, map01100, map01110, map01120, map01130
KEGG modules M00135
Gene Ontology (54) GO:0000166, GO:0003674, GO:0003824, GO:0004029, GO:0004030, GO:0005488, GO:0005575, GO:0005622, GO:0005623, GO:0005737, GO:0005739, GO:0005759 +42 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.177 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 10 synonymous, 5 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
AldedhPF00171.28 9.7e-14522–483 Aldehyde dehydrogenase family

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0765c (oxidoreductase), high confidence from genomic context alone (score 958 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0765c oxidoreductase 958 958 ctx neighborhood:758 coexpression:835
Rv0770 oxidoreductase 957 956 ctx neighborhood:746 coexpression:832
Rv0761c adhB alcohol dehydrogenase B 954 952 ctx neighborhood:438 database:900
Rv0766c cyp123 cytochrome P450 Cyp123 953 952 ctx neighborhood:758 coexpression:807
Rv3667 acs acetyl-CoAsynthetase 950 947 coexpression:418 database:900
Rv0764c cyp51 lanosterol 14-alpha demethylase 943 943 ctx neighborhood:758 coexpression:736
Rv0771 4-carboxymuconolactone decarboxylase 941 939 ctx neighborhood:746 coexpression:736
Rv2589 gabT 4-aminobutyrate aminotransferase 937 932 database:900
Rv0753c mmsA methylmalonate-semialdehyde dehydrogenase 929 927 database:900
Rv0409 ackA acetate kinase 930 922 database:900
Rv0860 fadB fatty oxidation protein FadB 925 921 database:900
Rv1530 adh alcohol dehydrogenase 924 920 database:900
Rv0162c adhE1 zinc-type alcohol dehydrogenase subunit E 924 920 database:900
Rv3432c gadB glutamate decarboxylase GadB 921 916 database:900
Rv1862 adhA alcohol dehydrogenase A 918 914 database:900

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: aldehyde dehydrogenase AldA
  • MTBC0 PGAP product: aldehyde dehydrogenase
  • Pfam (hmmscan --cut_ga): Aldedh PF00171.28 (E=1e-144)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215282.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Aldedh (PF00171.28)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1012
  • Curated reference: UniProt I6X9R9 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 93 functional partner(s); context anchor Rv0765c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000817|Rv0768|aldA
MALWGDGISALLIDGKLSDGRAGTFPTVNPATEEVLGVAADADAEDMGRAIEAARRAFDSTDWSRNTELRVRCVRQLRDAMQQHVEELRELTISEVGAPRMLTASAQLEGPVGDLSFAADTAESYPWKQDLGEASPLGIATRRTLAREAVGVVGAITPWNFPHQINLAKLGPALAAGNTVVLKPAPDTPWCAAALGEIIVEHTDFPPGVVNIVTSSSHALGALLAKDPRVDMISFTGSTATGRAVMADAAATIKKVFLELGGKSAFVVLDDADLAAASAVSAFSACMHAGQGCAITTRLVVPRARYEEAVAIAAATMSSIRPGDPNDPGTVCGPLISARQRDRVQGYLDLAVAEGGRFACGGARPADREVGFYIEPTVIAGLTNDARVAREEIFGPVLTVIAHDGDDDAVRIANDSPYGLSGTVYGADPQRAARIASRLRVGTVNVNGGVWYCADAPFGGYKQSGIGREMGLLGFEEYLEAKLIATAAN