Rv0769 Family assigned · medium auto-curated

H37Rv Rv0769 · MTBC0 mtbc0_000818 · 248 aa · 866836–867582 (+) · RefSeq NP_215283.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)oxidoreductase
MTBC0 PGAP re-annotationSDR family oxidoreductase
Revised (this work)SDR family oxidoreductase. Pfam: adh_short (PF00106.32), KR (PF08659.17), adh_short_C2 (PF13561.13).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WGQ9 SwissProt · reviewed · Evidence at protein level
UniProt nameUncharacterized oxidoreductase Rv0769
EC (curated) EC 1.-.-.-

UniProt still lists this protein as Uncharacterized oxidoreductase Rv0769; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category I Lipid transport and metabolism
Q Secondary metabolites biosynthesis, transport and catabolism
eggNOG descriptionDehydrogenase
Orthologous groupCOG1028
EC number EC 1.1.1.100
KEGG orthology K00059
KEGG pathways map00061, map00333, map00780, map01040, map01100, map01130, map01212
KEGG modules M00083, M00572
Gene Ontology (6) GO:0005575, GO:0005623, GO:0005886, GO:0016020, GO:0044464, GO:0071944

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.443 · purifying
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
adh_shortPF00106.32 7.4e-515–196 short chain dehydrogenase
KRPF08659.17 1.2e-058–92 KR domain
adh_short_C2PF13561.13 2.3e-6511–243 Enoyl-(Acyl carrier protein) reductase

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: htdY (3-hydroxyacyl-thioester dehydratase HtdY), high confidence from genomic context alone (score 930 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv2524c fas fatty acid synthase 981 979 coexpression:506 experimental:475 database:918
Rv2243 fabD malonyl CoA-acyl carrier protein transacylase 945 943 database:900
Rv3389c htdY 3-hydroxyacyl-thioester dehydratase HtdY 931 930 ctx fusion:899
Rv3559c oxidoreductase 924 925 database:900
Rv1350 fabG2 3-oxoacyl-ACP reductase FabG 917 917 database:900
Rv3538 dehydrogenase 916 916 ctx fusion:878
Rv0242c fabG4 3-oxoacyl-ACP reductase FabG 913 914 database:900
Rv0649 fabD2 malonyl CoA-acyl carrier protein transacylase 900 900 database:900
Rv0768 aldA aldehyde dehydrogenase AldA 872 867 ctx neighborhood:836
Rv0771 4-carboxymuconolactone decarboxylase 831 825 ctx neighborhood:773
Rv0762c hyp hypothetical protein 811 811 ctx neighborhood:731
Rv0765c oxidoreductase 808 808 ctx neighborhood:731
Rv0764c cyp51 lanosterol 14-alpha demethylase 798 798 ctx neighborhood:731
Rv0772 purD phosphoribosylamine--glycine ligase 802 796 ctx neighborhood:768
Rv0766c cyp123 cytochrome P450 Cyp123 795 795 ctx neighborhood:731

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: oxidoreductase
  • MTBC0 PGAP product: SDR family oxidoreductase
  • Pfam (hmmscan --cut_ga): adh_short PF00106.32 (E=7e-51), KR PF08659.17 (E=1e-05), adh_short_C2 PF13561.13 (E=2e-65)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215283.1)
  • Domains: Pfam-A via hmmscan --cut_ga — adh_short (PF00106.32), KR (PF08659.17), adh_short_C2 (PF13561.13)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1028
  • Curated reference: UniProt P9WGQ9 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 85 functional partner(s); context anchor htdY
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000818|Rv0769|
MFDSKVAIVTGAAQGIGQAYAQALAREGASVVVADINADGAAAVAKQIVADGGTAIHVPVDVSDEDSAKAMVDRAVGAFGGIDYLVNNAAIYGGMKLDLLLTVPLDYYKKFMSVNHDGVLVCTRAVYKHMAKRGGGAIVNQSSTAAWLYSNFYGLAKVGVNGLTQQLARELGGMKIRINAIAPGPIDTEATRTVTPAELVKNMVQTIPLSRMGTPEDLVGMCLFLLSDSASWITGQIFNVDGGQIIRS