Rv0204c Family assigned · medium auto-curated

H37Rv Rv0204c · MTBC0 mtbc0_000218 · 412 aa · 242328–243566 (-) · RefSeq NP_214718.2

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)transmembrane protein
MTBC0 PGAP re-annotationTIGR00374 family protein
Revised (this work)TIGR00374 family protein. Pfam: LPG_synthase_TM (PF03706.19).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6Y748 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable conserved transmembrane protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionintegral membrane protein
Orthologous groupCOG0392
KEGG orthology K20468

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.542 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
LPG_synthase_TMPF03706.19 4.5e-4044–346 Lysylphosphatidylglycerol synthase TM region

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0205 (transmembrane protein), high confidence from genomic context alone (score 774 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0205 transmembrane protein 972 774 ctx neighborhood:767 textmining:883
Rv0128 transmembrane protein 716 717 ctx cooccurence:706
Rv0955 integral membrane protein 680 680 ctx cooccurence:671
Rv0236c aftD alpha-(1->3)-arabinofuranosyltransferase 739 666 ctx cooccurence:616
Rv1978 hyp hypothetical protein 643 643 ctx cooccurence:631
Rv3912 rsmA anti-sigma-M factor RsmA 616 617 ctx cooccurence:604
Rv0048c membrane protein 582 583 ctx cooccurence:580
Rv2248 hyp hypothetical protein 582 583 ctx cooccurence:581
Rv2181 alpha-(1-2)-phosphatidylinositol mannoside mannosyltransferase 575 575 ctx cooccurence:567
Rv0256c PPE2 PPE family protein PPE2 567 567 ctx cooccurence:562
Rv3604c transmembrane protein 567 567 ctx cooccurence:561
Rv1863c integral membrane protein 566 567 ctx cooccurence:543
Rv2942 mmpL7 transmembrane transport protein MmpL7 563 563 ctx cooccurence:546
Rv3779 transmembrane protein 561 561 ctx cooccurence:551
Rv2687c antibiotic ABC transporter permease 535 535 ctx cooccurence:530

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: transmembrane protein
  • MTBC0 PGAP product: TIGR00374 family protein
  • Pfam (hmmscan --cut_ga): LPG_synthase_TM PF03706.19 (E=5e-40)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214718.2)
  • Domains: Pfam-A via hmmscan --cut_ga — LPG_synthase_TM (PF03706.19)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0392
  • Curated reference: UniProt I6Y748 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 50 functional partner(s); context anchor Rv0205
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000218|Rv0204c|
MSHDAPARNLRQRVGALPRTRVGAPPAEGVPPRGKYWWLRWAVLAIVAIVLAIEVALGWDQLAKAWVSLYRAKWWWLLAAVAAAGASMHSFAQIQRTLLKSAGVHVKQWRSEAAFYAANSLSTTLPGGPVLSATFLLRQQRIWGASTVVASWQLVMSGVLQAVGLALLGLGGAFFLGAKNNPFSLLFTLGGFVTLLLLAQAVASRPELIEGIGRRVLSWANSVRGRPADAGLPKWRETLMQLESVSLGRRDLGVAFGWSLFNWIADVACLGFAAYAAGDHASVGGLAVAYAAARAVGTIPLMPGGLLVVEAVLVPGLVSSGMPLPSAISAMLIYRLISWLLIAAIGWVVFFFMFRTESTADSDNDRDPPTDPNLRLVIQPQGTPCDDPVETTPQGPAPTPDLRPEGGETPPR