Rv0195 Resolved · high auto-curated

H37Rv Rv0195 · MTBC0 mtbc0_000209 · 211 aa · 231249–231884 (+) · RefSeq NP_214709.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)two component transcriptional regulator
MTBC0 PGAP re-annotationresponse regulator transcription factor
Revised (this work)Response regulator transcription factor. Pfam: GerE (PF00196.26).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O53646 TrEMBL · unreviewed · Predicted
UniProt namePossible two component transcriptional regulatory protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category K Transcription
eggNOG descriptionBacterial regulatory proteins, luxR family
Orthologous groupCOG2197

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains) pseudogene candidate

pN/pS 0.712 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 4 missense, 0 nonsense, 1 frameshift
Disruption 1 distinct premature-stop/frameshift site(s); most common in 6.00% of strains (8709) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
GerEPF00196.26 2.4e-12150–205 Bacterial regulatory proteins, luxR family

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: narS (sensor histidine kinase NarS), high confidence from genomic context alone (score 818 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0194 multidrug ABC transporter ATPase/permease 866 831 coexpression:826
Rv0845 narS sensor histidine kinase NarS 839 818 ctx cooccurence:662
Rv0193c hyp hypothetical protein 903 732 coexpression:731 textmining:655
Rv0844c narL nitrate/nitrite response transcriptional regulator NarL 712 713 ctx cooccurence:708
Rv3133c devR two component transcriptional regulator DevR 780 704 ctx cooccurence:703
Rv2027c dosT two component sensor histidine kinase DosT 749 650
Rv3132c devS two component sensor histidine kinase DevS 725 642
Rv0197 oxidoreductase 542 527 ctx neighborhood:526
Rv0196 HTH-type transcriptional regulator 528 527 ctx neighborhood:526
Rv1674c transcriptional regulator 509 478 coexpression:476
Rv3840 transcriptional regulator 433 426
Rv0601c two component sensor kinase HK2 417 395
Rv1675c cmr HTH-type transcriptional regulator Cmr 424 394
Rv1189 sigI ECF RNA polymerase sigma factor SigI 408 387
Rv0494 HTH-type transcriptional regulator 406 383

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: two component transcriptional regulator
  • MTBC0 PGAP product: response regulator transcription factor
  • Pfam (hmmscan --cut_ga): GerE PF00196.26 (E=2e-12)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214709.1)
  • Domains: Pfam-A via hmmscan --cut_ga — GerE (PF00196.26)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2197
  • Curated reference: UniProt O53646 (TrEMBL, unreviewed; Predicted)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 28 functional partner(s); context anchor narS
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000209|Rv0195|
MAPVNVISVAVVASDPLTRDGALARLSSHRELDVRAWQAGCETSVLLVLATTITAPLLCQIEDVQKDGPSHAPKLVVVADEFSAEQVFRMIKLGLTGLLYRSQSTFDCIVETIRLSAEGRLRLPERVQRYLVGRIKSTPTAEPDTPCAAALAEREVAVLRLLADGLSTHQVAVQLNYCERTIKNIVHDIVTRLKLRNRTHAVAHALRAGLI