Rv0205 Family assigned · medium auto-curated

H37Rv Rv0205 · MTBC0 mtbc0_000219 · 367 aa · 243736–244839 (+) · RefSeq NP_214719.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)transmembrane protein
MTBC0 PGAP re-annotationAI-2E family transporter
Revised (this work)AI-2E family transporter. Pfam: AI-2E_transport (PF01594.23).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WFM5 SwissProt · reviewed · Evidence at protein level
UniProt namePutative transport protein Rv0205

UniProt still lists this protein as Putative transport protein Rv0205; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionAI-2E family transporter
Orthologous groupCOG0628
KEGG orthology K20469
Gene Ontology (20) GO:0003674, GO:0005215, GO:0005575, GO:0005623, GO:0005886, GO:0005887, GO:0006810, GO:0008150, GO:0016020, GO:0016021, GO:0031224, GO:0031226 +8 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains) pseudogene candidate

pN/pS 0.772 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 6 missense, 0 nonsense, 1 frameshift
Disruption 1 distinct premature-stop/frameshift site(s); most common in 4.91% of strains (7135) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
AI-2E_transportPF01594.23 1.2e-7925–348 AI-2E family transporter

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0204c (transmembrane protein), high confidence from genomic context alone (score 774 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0204c transmembrane protein 972 774 ctx neighborhood:767 textmining:883
Rv0206c mmpL3 transmembrane transport protein MmpL3 498 145 textmining:437
Rv0202c mmpL11 transmembrane transport protein MmpL11 669 144 textmining:630
Rv0207c hyp hypothetical protein 493 67 textmining:479
Rv0201c hyp hypothetical protein 807 64 textmining:803
Rv0200 transmembrane protein 555 52 textmining:550
Rv0208c trmB tRNA (guanine-N(7)-)-methyltransferase 651 51 textmining:648
Rv1972 Mce associated membrane protein 657 41 textmining:657
Rv0177 Mce associated protein 550 41 textmining:550
Rv3444c esxT ESAT-6 like protein EsxT 510 41 textmining:510

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: transmembrane protein
  • MTBC0 PGAP product: AI-2E family transporter
  • Pfam (hmmscan --cut_ga): AI-2E_transport PF01594.23 (E=1e-79)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214719.1)
  • Domains: Pfam-A via hmmscan --cut_ga — AI-2E_transport (PF01594.23)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0628
  • Curated reference: UniProt P9WFM5 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 10 functional partner(s); context anchor Rv0204c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000219|Rv0205|
MSASLDDASVAPLVRKTAAWAWRFLVILAAMVALLWVLNKFEVIVVPVLLALMLSALLVPPVDWLDSRGLPRAVAVTLVLLSGFAVLGGILTFVVSQFIAGLPHLVTEVERSIDSARRWLIEGPAHLRGEQIDNAGNAAIEALRNNQAKLTSGALSTAATITELVTAAVLVLFTLIFFLYGGRSIWQYVTKAFPASVRDRVRAAGRAGYASLIGYARATFLVALTDAAGVGAGLAVMGVPLALPLASLVFFGAFIPLIGAVVAGFLAVVVALLAKGIGYALITVGLLIAVNQLEAHLLQPLVMGRAVSIHPLAVVLAIAAGGVLAGVVGALLAVPTVAFFNNAVQVLLGGNPFADVADVSSDHLTEV