pirG Resolved · medium auto-curated

H37Rv Rv3810 · MTBC0 mtbc0_004038 · 284 aa · 4297849–4298703 (+) · RefSeq NP_218327.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)cell surface protein
MTBC0 PGAP re-annotationexported repetitive protein
Revised (this work)Exported repetitive protein.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WIQ7 SwissProt · reviewed · Evidence at protein level
UniProt nameExported repetitive protein
Curated functionSurface-exposed protein required for multiplication and intracellular growth.

Functional vocabulary (eggNOG-mapper, orthology transfer)

Preferred nameerp
Orthologous group2EIXY
Gene Ontology (12) GO:0005575, GO:0005576, GO:0008150, GO:0009405, GO:0040007, GO:0044110, GO:0044116, GO:0044117, GO:0044119, GO:0044403, GO:0044419, GO:0051704

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 1.171 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: eccD3 (ESX-3 secretion system protein EccD), high confidence from genomic context alone (score 769 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv2721c hyp hypothetical protein 918 918 ctx cooccurence:709 coexpression:730
Rv0312 hyp hypothetical protein 865 860 ctx cooccurence:452 coexpression:754
Rv0129c fbpC diacylglycerol acyltransferase/mycolyltransferase Ag85C 853 852 coexpression:765
Rv1477 ripA peptidoglycan endopeptidase RipA 843 843 coexpression:801
Rv1158c hyp hypothetical protein 831 824 coexpression:772
Rv0040c mtc28 hyp hypothetical protein 812 812 coexpression:812
Rv1566c ripD hyp hypothetical protein 809 802 coexpression:802
Rv3811 hyp hypothetical protein 787 779 ctx neighborhood:714
Rv1157c hyp hypothetical protein 775 774 coexpression:733
Rv3707c hyp hypothetical protein 770 770 ctx cooccurence:769
Rv0290 eccD3 ESX-3 secretion system protein EccD 769 769 ctx cooccurence:766
Rv0479c membrane protein 767 767 ctx cooccurence:759
Rv3446c hyp hypothetical protein 770 762 ctx cooccurence:761
Rv2743c hyp hypothetical protein 762 762 ctx cooccurence:760
Rv0256c PPE2 PPE family protein PPE2 761 762 ctx cooccurence:758

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: cell surface protein
  • MTBC0 PGAP product: exported repetitive protein
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218327.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 2EIXY
  • Curated reference: UniProt P9WIQ7 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 162 functional partner(s); context anchor eccD3
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_004038|Rv3810|pirG
MPNRRRRKLSTAMSAVAALAVASPCAYFLVYESTETTERPEHHEFKQAAVLTDLPGELMSALSQGLSQFGINIPPVPSLTGSGDASTGLTGPGLTSPGLTSPGLTSPGLTDPALTSPGLTPTLPGSLAAPGTTLAPTPGVGANPALTNPALTSPTGATPGLTSPTGLDPALGGANEIPITTPVGLDPGADGTYPILGDPTLGTIPSSPATTSTGGGGLVNDVMQVANELGASQAIDLLKGVLMPSIMQAVQNGGAAAPAASPPVPPIPAAAAVPPTDPITVPVA