Rv3679 Family assigned · medium auto-curated

H37Rv Rv3679 · MTBC0 mtbc0_003899 · 340 aa · 4142606–4143628 (+) · RefSeq NP_218196.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)anion transporter ATPase
MTBC0 PGAP re-annotationArsA family ATPase
Revised (this work)ArsA family ATPase. Pfam: ArsA_ATPase (PF02374.22), CbiA (PF01656.30).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WKX5 SwissProt · reviewed · Evidence at protein level
UniProt nameATPase BagA
EC (curated) EC 3.6.1.-
Curated functionComponent of the heterodimeric BagAB ATPase complex, whose two subunits are actively involved in ATP hydrolysis. The ATPase activity is required to mediate resistance against nitric oxide (NO) and elevated levels of glycerol.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category P Inorganic ion transport and metabolism
Preferred namearsA_2
eggNOG descriptionPFAM Anion-transporting ATPase
Orthologous groupCOG0003
Gene Ontology (8) GO:0005575, GO:0005623, GO:0005886, GO:0008150, GO:0016020, GO:0040007, GO:0044464, GO:0071944

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.214 · purifying
Polymorphic sites (≥ 0.1% of strains) 5 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
ArsA_ATPasePF02374.22 2.1e-1522–181 Anion-transporting ATPase
CbiAPF01656.30 1.6e-0726–241 CobQ/CobB/MinD/ParA nucleotide binding domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3680 (anion transporter ATPase), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3680 anion transporter ATPase 999 1000 ctx neighborhood:882 experimental:999 database:900
Rv3678A hyp hypothetical protein 785 785 ctx neighborhood:783
Rv3678c hyp hypothetical protein 781 781 ctx neighborhood:774
Rv3677c beta lactamase 783 776 ctx neighborhood:774
Rv0487 hyp hypothetical protein 759 759 ctx cooccurence:758
Rv2185c TB16.3 hyp hypothetical protein 720 720 ctx cooccurence:708
Rv0857 hyp hypothetical protein 715 716 ctx cooccurence:715
Rv0854 hyp hypothetical protein 697 697 ctx cooccurence:686
Rv0244c fadE5 acyl-CoA dehydrogenase FadE5 684 685 ctx cooccurence:682
Rv0474 HTH-type transcriptional regulator 665 665 ctx cooccurence:663
Rv1467c fadE15 acyl-CoA dehydrogenase 651 652 ctx cooccurence:647
Rv0501 galE2 UDP-glucose 4-epimerase GalE 630 630 ctx cooccurence:627
Rv0502 hyp hypothetical protein 624 624 ctx cooccurence:619
Rv3662c hyp hypothetical protein 585 585 ctx cooccurence:582
Rv1691 hyp hypothetical protein 578 556 database:410

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: anion transporter ATPase
  • MTBC0 PGAP product: ArsA family ATPase
  • Pfam (hmmscan --cut_ga): ArsA_ATPase PF02374.22 (E=2e-15), CbiA PF01656.30 (E=2e-07)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218196.1)
  • Domains: Pfam-A via hmmscan --cut_ga — ArsA_ATPase (PF02374.22), CbiA (PF01656.30)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0003
  • Curated reference: UniProt P9WKX5 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 29 functional partner(s); context anchor Rv3680
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003899|Rv3679|
MVATTSSGGSSVGWPSRLSGVRLHLVTGKGGTGKSTIAAALALTLAAGGRKVLLVEVEGRQGIAQLFDVPPLPYQELKIATAERGGQVNALAIDIEAAFLEYLDMFYNLGIAGRAMRRIGAVEFATTIAPGLRDVLLTGKIKETVVRLDKNKLPVYDAIVVDAPPTGRIARFLDVTKAVSDLAKGGPVHAQSEGVVKLLHSNQTAIHLVTLLEALPVQETLEAIEELAQMELPIGSVIVNRNIPAHLEPQDLAKAAEGEVDADSVRAGLLTAGVKLPDADFAGLLTETIQHATRITARAEIAQQLDALQVPRLELPTVSDGVDLGSLYELSESLAQQGVR