Rv2844 Family assigned · low auto-curated

H37Rv Rv2844 · MTBC0 mtbc0_003023 · 162 aa · 3171379–3171867 (+) · RefSeq NP_217360.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationferritin-like domain-containing protein
Revised (this work)Ferritin-like domain-containing protein.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6Y1V1 TrEMBL · unreviewed · Evidence at protein level
UniProt nameConserved alanine rich protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionDomain of unknown function (DUF4439)
Orthologous group2EA6A

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.0 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 0 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
DUF4439PF14530.13 7.4e-4625–161 Domain of unknown function (DUF4439)

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: nusA (transcription termination/antitermination protein NusA), high confidence from genomic context alone (score 762 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv2843 hyp hypothetical protein 913 913 ctx neighborhood:882
Rv2841c nusA transcription termination/antitermination protein NusA 762 762 ctx neighborhood:760
Rv2842c rimP ribosome maturation factor RimP 761 762 ctx neighborhood:760
Rv3118 sseC1 hyp hypothetical protein 727 728 ctx cooccurence:726
Rv0814c sseC2 hyp hypothetical protein 726 727 ctx cooccurence:726
Rv3605c hyp hypothetical protein 660 660 ctx cooccurence:660
Rv3231c hyp hypothetical protein 623 624 ctx cooccurence:615
Rv2728c hyp hypothetical protein 619 619 ctx cooccurence:618
Rv1332 transcriptional regulator 609 609 ctx cooccurence:608
Rv2468c hyp hypothetical protein 598 598 ctx cooccurence:589
Rv0010c membrane protein 569 569 ctx cooccurence:568
Rv3839 hyp hypothetical protein 561 561 ctx cooccurence:559
Rv1486c hyp hypothetical protein 551 551 ctx cooccurence:550
Rv1473A transcriptional regulator 546 546 ctx cooccurence:541
Rv2712c hyp hypothetical protein 538 538 ctx cooccurence:537

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: ferritin-like domain-containing protein
  • Pfam (hmmscan --cut_ga): DUF4439 PF14530.13 (E=7e-46)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217360.1)
  • Domains: Pfam-A via hmmscan --cut_ga — DUF4439 (PF14530.13)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 2EA6A
  • Curated reference: UniProt I6Y1V1 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 35 functional partner(s); context anchor nusA
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003023|Rv2844|
MTSSEPAHGATPKRSPSEGSADNAALCDALAVEHATIYGYGIVSALSPPGVNFLVADALKQHRHRRDDVIVMLSARGVTAPIAAAGYQLPMQVSSAADAARLAVRMENDGATAWRAVVEHAETADDRVFASTALTESAVMATRWNRVLGAWPITAAFPGGDE