Rv1505c Resolved · high auto-curated

H37Rv Rv1505c · MTBC0 mtbc0_001609 · 221 aa · 1705086–1705751 (-) · RefSeq NP_216021.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationacetyltransferase
Revised (this work)Acetyltransferase. Pfam: Hexapep (PF00132.31), Hexapep_2 (PF14602.13).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P71784 TrEMBL · unreviewed · Evidence at protein level
UniProt nameTransferase

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionHexapeptide repeat of succinyl-transferase
Orthologous groupCOG0110

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.793 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 5 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
HexapepPF00132.31 1.5e-07160–193 Bacterial transferase hexapeptide (six repeats)
Hexapep_2PF14602.13 6.3e-06161–191 Hexapeptide repeat of succinyl-transferase

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv1504c (Rv1504c, (MTCY277.26c), len: 199 aa. Conserved hypothetical protein, similar to N-terminal region of P27833|RFFA_ECOLI lipopolysaccharide bi), high confidence from genomic context alone (score 977 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv1504c Rv1504c, (MTCY277.26c), len: 199 aa. Conserved hypothetical protein, similar to N-terminal region of P27833|RFFA_ECOLI lipopolysaccharide bi 978 977 ctx neighborhood:516 cooccurence:651 coexpression:875
Rv1506c hyp hypothetical protein 974 947 ctx neighborhood:781 cooccurence:626 coexpression:409 textmining:527
Rv1503c Rv1503c, (MTCY277.25c), len: 182 aa. Conserved hypothetical protein, similar to C-terminal region of P27833|RFFA_ECOLI lipopolysaccharide bi 948 906 ctx neighborhood:516 cooccurence:608 coexpression:545 textmining:471
Rv1507c hyp hypothetical protein 887 864 ctx neighborhood:443 cooccurence:752
Rv1502 hyp hypothetical protein 815 798 ctx cooccurence:773
Rv3402c hyp hypothetical protein 756 741 ctx cooccurence:570
Rv1501 hyp hypothetical protein 445 445 ctx cooccurence:400
Rv1500 pimF glycosyltransferase 453 428
Rv1519 hyp hypothetical protein 457 424
Rv1570 bioD ATP-dependent dethiobiotin synthetase BioD 430 401 coexpression:401
Rv3556c fadA6 acetyl-CoA acetyltransferase FadA 573 242 textmining:461
Rv3523 ltp3 lipid carrier protein 568 235 textmining:459
Rv3546 fadA5 acetyl-CoA acetyltransferase FadA 438 230
Rv3225c GCN5-like N-acetyltransferase 509 225
Rv1527c pks5 polyketide synthase 578 183 textmining:506

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: acetyltransferase
  • Pfam (hmmscan --cut_ga): Hexapep PF00132.31 (E=1e-07), Hexapep_2 PF14602.13 (E=6e-06)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216021.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Hexapep (PF00132.31), Hexapep_2 (PF14602.13)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0110
  • Curated reference: UniProt P71784 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 21 functional partner(s); context anchor Rv1504c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001609|Rv1505c|
MTKPLVIFGSGDIAQLAHYYFTRDSEYEVVAFTVDRDYASVSEFCGLPLVAFDEVAQRFPPESHAMFVALAYAKLNGVRKEKYLAAKALGYELASYVSSHATVLNDGRIGENVFLLEDNTIQPFVSIGNNVTLWSGNHIGHHSTIHDHCFLASHIVVSGGVVIEEQSFIGVNATLRDHITIGSRCVVGAGALLLGDADADGVYIGTKTERRPVPSTELRKI