ctpE Resolved · high auto-curated

H37Rv Rv0908 · MTBC0 mtbc0_000962 · 797 aa · 1014946–1017339 (+) · RefSeq NP_215423.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)metal cation transporter ATPase E
MTBC0 PGAP re-annotationcation-translocating P-type ATPase
Revised (this work)Cation-translocating P-type ATPase. Pfam: E1-E2_ATPase (PF00122.26), Hydrolase (PF00702.33), Hydrolase_3 (PF08282.19).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WPT1 SwissProt · reviewed · Evidence at protein level
UniProt nameCalcium-transporting ATPase CtpE
EC (curated) EC 7.2.2.10
Curated functionP-type ATPase involved in specific uptake of calcium.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category P Inorganic ion transport and metabolism
Preferred namectpE
eggNOG descriptionATPase, P-type (transporting), HAD superfamily, subfamily IC
Orthologous groupCOG0474
KEGG orthology K12952
Gene Ontology (64) GO:0003674, GO:0003824, GO:0005215, GO:0005388, GO:0005575, GO:0005622, GO:0005623, GO:0005886, GO:0005887, GO:0006810, GO:0006811, GO:0006812 +52 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.205 · purifying
Polymorphic sites (≥ 0.1% of strains) 7 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
E1-E2_ATPasePF00122.26 1.8e-23104–196 P-type ATPase actuator domain
HydrolasePF00702.33 5.7e-16295–548 haloacid dehalogenase-like hydrolase
Hydrolase_3PF08282.19 8.5e-05520–571 haloacid dehalogenase-like hydrolase

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: echA6 (enoyl-CoA hydratase EchA6), medium confidence from genomic context alone (score 685 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0907 hyp hypothetical protein 939 903 ctx neighborhood:881
Rv0906 hyp hypothetical protein 789 790 ctx neighborhood:787
Rv0905 echA6 enoyl-CoA hydratase EchA6 685 685 ctx neighborhood:682
Rv1747 ABC transporter ATP-binding protein/permease 663 641 coexpression:421
Rv1305 atpE ATP synthase subunit C 630 608 database:551
Rv0909 antitoxin 576 576 ctx neighborhood:544
Rv1488 hyp hypothetical protein 558 533
Rv3008 hyp hypothetical protein 546 518 coexpression:410
Rv1811 mgtC Mg2+ transport P-type ATPase MgtC 545 517 coexpression:409
Rv0904c accD3 acetyl-CoAcarboxylase carboxyl transferase subunit beta 533 495 ctx neighborhood:493
Rv3005c hyp hypothetical protein 485 485
Rv2483c plsC bifunctional L-3-phosphoserine phosphatase/1-acyl-sn-glycerol-3-phosphate acyltransferase 506 484 database:451
Rv3486 hyp hypothetical protein 483 483
Rv3064c integral membrane protein 481 481
Rv0924c mntH divalent metal cation transporter MntH 645 480

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: metal cation transporter ATPase E
  • MTBC0 PGAP product: cation-translocating P-type ATPase
  • Pfam (hmmscan --cut_ga): E1-E2_ATPase PF00122.26 (E=2e-23), Hydrolase PF00702.33 (E=6e-16), Hydrolase_3 PF08282.19 (E=9e-05)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215423.1)
  • Domains: Pfam-A via hmmscan --cut_ga — E1-E2_ATPase (PF00122.26), Hydrolase (PF00702.33), Hydrolase_3 (PF08282.19)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0474
  • Curated reference: UniProt P9WPT1 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 49 functional partner(s); context anchor echA6
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000962|Rv0908|ctpE
MTRSASATAGLTDAEVAQRVAEGKSNDIPERVTRTVGQIVRANVFTRINAILGVLLLIVLATGSLINGMFGLLIIANSVIGMVQEIRAKQTLDKLAIIGQAKPLVRRQSGTRTRSTNEVVLDDIIELGPGDQVVVDGEVVEEENLEIDESLLTGEADPIAKDAGDTVMSGSFVVSGAGAYRATKVGSEAYAAKLAAEASKFTLVKSELRNGINRILQFITYLLVPAGLLTIYTQLFTTHVGWRESVLRMVGALVPMVPEGLVLMTSIAFAVGVVRLGQRQCLVQELPAIEGLARVDVVCADKTGTLTESGMRVCEVEELDGAGRQESVADVLAALAAADARPNASMQAIAEAFHSPPGWVVAANAPFKSATKWSGVSFRDHGNWVIGAPDVLLDPASVAARQAERIGAQGLRVLLLAAGSVAVDHAQAPGQVTPVALVVLEQKVRPDARETLDYFAVQNVSVKVISGDNAVSVGAVADRLGLHGEAMDARALPTGREELADTLDSYTSFGRVRPDQKRAIVHALQSHGHTVAMTGDGVNDVLALKDADIGVAMGSGSPASRAVAQIVLLNNRFATLPHVVGEGRRVIGNIERVANLFLTKTVYSVLLALLVGIECLIAIPLRRDPLLFPFQPIHVTIAAWFTIGIPAFILSLAPNNERAYPGFVRRVMTSAVPFGLVIGVATFVTYLAAYQGRYASWQEQEQASTAALITLLMTALWVLAVIARPYQWWRLALVLASGLAYVVIFSLPLAREKFLLDASNLATTSIALAVGVVGAATIEAMWWIRSRMLGVKPRVWR