Rv1747 Family assigned · medium auto-curated

H37Rv Rv1747 · MTBC0 mtbc0_001860 · 865 aa · 1985702–1988299 (+) · RefSeq NP_216263.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)ABC transporter ATP-binding protein/permease
MTBC0 PGAP re-annotationATP-binding cassette domain-containing protein
Revised (this work)ATP-binding cassette domain-containing protein. Pfam: Yop-YscD_cpl (PF16697.11), FHA (PF00498.32), ABC_tran (PF00005.34), AAA_21 (PF13304.13), ABC2_membrane (PF01061.30).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O65934 SwissProt · reviewed · Evidence at protein level
UniProt nameABC transporter ATP-binding/permease protein Rv1747
EC (curated) EC 7.-.-.-
Curated functionInvolved in the translocation of an unknown substrate across the membrane. Transmembrane domains (TMD) form a pore in the membrane and the ATP-binding domain (NBD) is responsible for energy generation. Required for virulence.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category V Defense mechanisms
eggNOG descriptionABC transporter
Orthologous groupCOG0842
KEGG orthology K01990, K21397
KEGG modules M00254
Gene Ontology (37) GO:0003674, GO:0003824, GO:0005215, GO:0005575, GO:0005618, GO:0005622, GO:0005623, GO:0005737, GO:0005829, GO:0005886, GO:0006810, GO:0008150 +25 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.082 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 9 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Yop-YscD_cplPF16697.11 3.2e-10224–301 Inner membrane component of T3SS, cytoplasmic domain
FHAPF00498.32 2.9e-17230–293 FHA domain
ABC_tranPF00005.34 1.8e-32335–482 ABC transporter
AAA_21PF13304.13 1.2e-07437–514 AAA domain, putative AbiEii toxin, Type IV TA system
ABC2_membranePF01061.30 1.3e-19596–810 ABC-2 type transporter

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: pknF (serine/threonine-protein kinase PknF), high confidence from genomic context alone (score 739 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1248c kgd multifunctional 2-oxoglutarate dehydrogenase E1 component /2-oxoglutarate dehydrogenase dihydrolipoyllysine-residue succinyltransferase 989 985 experimental:984
Rv0014c pknB serine/threonine-protein kinase PknB 945 801 experimental:711 textmining:736
Rv1746 pknF serine/threonine-protein kinase PknF 952 739 ctx neighborhood:617 textmining:826
Rv1364c sigma factor regulatory protein 723 712 coexpression:647
Rv1278 hyp hypothetical protein 717 700 database:579
Rv1277 hyp hypothetical protein 703 692 database:579
Rv0434 hyp hypothetical protein 705 676 database:560
Rv3724A cut5a Rv3724A, (MTV025.072), len: 80 aa. Probable cut5a,truncated cutinase precursor, similar to N-terminal end of others e.g. Q9KK87 serine ester 662 649 database:530
Rv3724B cut5b Rv3724B, (MTV025.072), len: 187 aa. Probable cut5b,truncated cutinase, similar to C-terminal end of others e.g. Q9XB09|RVD2-RV1758 protein ( 662 649 database:530
Rv3483c hyp hypothetical protein 662 649 database:530
Rv2301 cut2 cutinase 661 648 database:530
Rv3451 cut3 cutinase 660 648 database:530
Rv1758 cut1 cutinase 660 647 database:530
Rv0908 ctpE metal cation transporter ATPase E 663 641 coexpression:421
Rv3132c devS two component sensor histidine kinase DevS 730 640 ctx neighborhood:497

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: ABC transporter ATP-binding protein/permease
  • MTBC0 PGAP product: ATP-binding cassette domain-containing protein
  • Pfam (hmmscan --cut_ga): Yop-YscD_cpl PF16697.11 (E=3e-10), FHA PF00498.32 (E=3e-17), ABC_tran PF00005.34 (E=2e-32), AAA_21 PF13304.13 (E=1e-07), ABC2_membrane PF01061.30 (E=1e-19)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216263.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Yop-YscD_cpl (PF16697.11), FHA (PF00498.32), ABC_tran (PF00005.34), AAA_21 (PF13304.13), ABC2_membrane (PF01061.30)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0842
  • Curated reference: UniProt O65934 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 171 functional partner(s); context anchor pknF
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001860|Rv1747|
MPMSQPAAPPVLTVRYEGSERTFAAGHDVVVGRDLRADVRVAHPLISRAHLLLRFDQGRWVAIDNGSLNGLYLNNRRVPVVDIYDAQRVHIGNPDGPALDFEVGRHRGSAGRPPQTTSIRLPNLSAGAWPTDGPPQTGTLGSGQLQQLPPATTRIPAAPPSGPQPRYPTGGQQLWPPSGPQRAPQIYRPPTAAPPPAGARGGTEAGNLATSMMKILRPGRLTGELPPGAVRIGRANDNDIVIPEVLASRHHATLVPTPGGTEIRDNRSINGTFVNGARVDAALLHDGDVVTIGNIDLVFADGTLARREENLLETRVGGLDVRGVTWTIDGDKTLLDGISLTARPGMLTAVIGPSGAGKSTLARLVAGYTHPTDGTVTFEGHNVHAEYASLRSRIGMVPQDDVVHGQLTVKHALMYAAELRLPPDTTKDDRTQVVARVLEELEMSKHIDTRVDKLSGGQRKRASVALELLTGPSLLILDEPTSGLDPALDRQVMTMLRQLADAGRVVLVVTHSLTYLDVCDQVLLLAPGGKTAFCGPPTQIGPVMGTTNWADIFSTVADDPDAAKARYLARTGPTPPPPPVEQPAELGDPAHTSLFRQFSTIARRQLRLIVSDRGYFVFLALLPFIMGALSMSVPGDVGFGFPNPMGDAPNEPGQILVLLNVGAVFMGTALTIRDLIGERAIFRREQAVGLSTTAYLIAKVCVYTVLAVVQSAIVTVIVLVGKGGPTQGAVALSKPDLELFVDVAVTCVASAMLGLALSAIAKSNEQIMPLLVVAVMSQLVFSGGMIPVTGRVPLDQMSWVTPARWGFAASAATVDLIKLVPGPLTPKDSHWHHTASAWWFDMAMLVALSVIYVGFVRWKIRLKAC