Rv0790c Family assigned · medium auto-curated

H37Rv Rv0790c · MTBC0 mtbc0_000841 · 242 aa · 888498–889226 (-) · RefSeq NP_215305.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationtransglutaminase-like domain-containing protein
Revised (this work)Transglutaminase-like domain-containing protein. Pfam: Transglut_core (PF01841.26).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6XWA9 TrEMBL · unreviewed · Evidence at protein level
UniProt nameTransglutaminase-like domain-containing protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category E Amino acid transport and metabolism
eggNOG descriptionTransglutaminase-like superfamily
Orthologous groupCOG1305

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.578 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 5 missense, 0 nonsense, 1 frameshift
Disruption 1 distinct premature-stop/frameshift site(s); most common in 0.26% of strains (374) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Transglut_corePF01841.26 2.9e-1829–152 Transglutaminase-like superfamily

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0792c (transcriptional regulator), high confidence from genomic context alone (score 888 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv0791c hyp hypothetical protein 998 987 ctx neighborhood:882 coexpression:860 textmining:870
Rv0792c transcriptional regulator 977 888 ctx neighborhood:818 textmining:806
Rv0793 monooxygenase 968 765 ctx neighborhood:609 textmining:870
Rv0789c hyp hypothetical protein 966 753 ctx neighborhood:750 textmining:870
Rv3387 transposase 587 587 ctx cooccurence:585
Rv1034c Probable transposase (fragment); Rv1034c, (MTCY10G2.15), len: 129 aa. Probable IS1560 transposase fragment, similar to part of Rv3387|E12023 550 550 ctx cooccurence:550
Rv2961 transposase 544 544 ctx cooccurence:543
Rv3386 transposase 518 518 ctx cooccurence:518
Rv0977 PE_PGRS16 PE-PGRS family protein PE_PGRS16 449 449 ctx cooccurence:445
Rv3081 hyp hypothetical protein 434 434 ctx cooccurence:429
Rv1749c integral membrane protein 429 429 ctx cooccurence:429
Rv3727 oxidoreductase 423 424 ctx cooccurence:422
Rv1452c PE_PGRS28 PE-PGRS family protein PE_PGRS28 417 418 ctx cooccurence:414
Rv0872c PE_PGRS15 PE-PGRS family protein PE_PGRS15 412 413 ctx cooccurence:409
Rv3531c hyp hypothetical protein 411 412 ctx cooccurence:408

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: transglutaminase-like domain-containing protein
  • Pfam (hmmscan --cut_ga): Transglut_core PF01841.26 (E=3e-18)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215305.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Transglut_core (PF01841.26)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1305
  • Curated reference: UniProt I6XWA9 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 19 functional partner(s); context anchor Rv0792c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000841|Rv0790c|
MTLANNGTGMDHFLTPTEYLDAGHPLVRTTAATLIRDAVSDTERVRRIYYYVRDVPYDVLASFRYLAQGHHRASDVIGHGVAFCMGKASSFVALCRAAGVPARIAFQTIDAPDKEFLSPQVRALWGGRTGRPFPWHSLGEAYLGRRWVKLDATIDAPTAARLGKPYRQEFDGATPIPTVEGTILRENGSYADYPSAVAQWYERIAQSVLKALQSTEVHALVAADEELWTGPPVELADATHRL