lprO Family assigned · medium auto-curated

H37Rv Rv0179c · MTBC0 mtbc0_000192 · 369 aa · 210051–211160 (-) · RefSeq NP_214693.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)lipoprotein LprO
MTBC0 PGAP re-annotationphosphodiester glycosidase family protein
Revised (this work)Phosphodiester glycosidase family protein. Pfam: NAGPA (PF09992.15).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O07423 TrEMBL · unreviewed · Evidence at protein level
UniProt namePossible lipoprotein LprO

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
Preferred namelprO
eggNOG descriptionPhosphodiester glycosidase
Orthologous group2F81I

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.192 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 7 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
NAGPAPF09992.15 7.8e-07250–345 Phosphodiester glycosidase

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0180c (transmembrane protein), medium confidence from genomic context alone (score 641 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv2145c wag31 cell wall synthesis protein Wag31 751 751 coexpression:751
Rv3414c sigD ECF RNA polymerase sigma factor SigD 730 730 coexpression:730
Rv3810 pirG cell surface protein 730 730 coexpression:730
Rv0180c transmembrane protein 764 641 ctx neighborhood:639
Rv0181c hyp hypothetical protein 782 564 ctx neighborhood:563 textmining:520
Rv1477 ripA peptidoglycan endopeptidase RipA 553 553 coexpression:553
Rv1987 chitinase 493 494 coexpression:494
Rv0182c sigG ECF RNA polymerase sigma factor SigG 573 487 ctx neighborhood:484
Rv2721c hyp hypothetical protein 474 474 coexpression:474
Rv3096 hyp hypothetical protein 441 441 coexpression:441
Rv2525c hyp hypothetical protein 427 193
Rv3495c lprN Mce family lipoprotein LprN 636 46 textmining:635
Rv1244 lpqZ lipoprotein LpqZ 602 46 textmining:601
Rv2518c ldtB L,D-transpeptidase LdtB 505 45 textmining:504
Rv2330c lppP lipoprotein LppP 616 41 textmining:617

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: lipoprotein LprO
  • MTBC0 PGAP product: phosphodiester glycosidase family protein
  • Pfam (hmmscan --cut_ga): NAGPA PF09992.15 (E=8e-07)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214693.1)
  • Domains: Pfam-A via hmmscan --cut_ga — NAGPA (PF09992.15)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 2F81I
  • Curated reference: UniProt O07423 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 16 functional partner(s); context anchor Rv0180c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000192|Rv0179c|lprO
MWIRAERVAVLTPTASLRRLTACYAALAVCAALACTTGQPAARAADGREMLAQAIATTRGSYLVYNFGGGHPMPLLNAGGHWYEMNNGGHLMIIKNASQRLSPHLLVDTHTGDQARCEHNPGARTGEGLWQASEIYPPLKAWQRMGRPTIAVNANFFDVRGQKGGSWRSTGCSSPLGAYVDNTRGQGRANQAVTGTVAYAGKQGLSGGNELWSSLTTMILPVGGAPYVLRPKSRQDYDLATPVIEDLLNKNARFVAVAGIGLLSPGNTGQLHDGGPSAARTALAYAKQKDEMYIFQGGNYTPDNIQDLFRGLGSDTAILLDGGGSSAIVLRRDTGGMWAGAGSPKGSCDTRQVLCDSHERALPSWLAFN