Rv0178 Resolved · medium auto-curated

H37Rv Rv0178 · MTBC0 mtbc0_000191 · 244 aa · 209286–210020 (+) · RefSeq NP_214692.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)Mce associated membrane protein
MTBC0 PGAP re-annotationMce associated membrane protein
Revised (this work)Mce associated membrane protein.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O07422 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable conserved Mce associated membrane protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

Orthologous group2BKHN
KEGG orthology K18481
KEGG modules M00670

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 1.747 · diversifying/relaxed
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 5 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0175 (Mce associated membrane protein), high confidence from genomic context alone (score 971 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0175 Mce associated membrane protein 992 971 ctx neighborhood:881 coexpression:770 textmining:751
Rv0176 Mce associated transmembrane protein 915 887 ctx neighborhood:801 coexpression:455
Rv0177 Mce associated protein 882 882 ctx neighborhood:882
Rv0174 mce1F Mce family protein Mce1F 924 840 ctx neighborhood:829 textmining:548
Rv0171 mce1C Mce family protein Mce1C 856 826 ctx neighborhood:821
Rv3301c phoY1 phosphate transport system transcriptional regulator PhoY 812 812 coexpression:812
Rv0014c pknB serine/threonine-protein kinase PknB 804 805 coexpression:798
Rv3692 moxR2 methanol dehydrogenase transcriptional regulator MoxR 802 803 coexpression:802
Rv3806c ubiA decaprenyl-phosphate phosphoribosyltransferase 773 773 coexpression:746
Rv2940c mas multifunctional mycocerosic acid synthase 769 742 experimental:470 database:508
Rv1663 pks17 polyketide synthase 750 741 experimental:458 database:508
Rv3825c pks2 phthioceranic/hydroxyphthioceranic acid synthase 765 737 experimental:470 database:508
Rv2933 ppsC phthiocerol synthesis polyketide synthase type I PpsC 764 737 experimental:470 database:508
Rv2048c pks12 polyketide synthase 764 737 experimental:470 database:508
Rv1527c pks5 polyketide synthase 763 735 experimental:470 database:508

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: Mce associated membrane protein
  • MTBC0 PGAP product: Mce associated membrane protein
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214692.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 2BKHN
  • Curated reference: UniProt O07422 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 273 functional partner(s); context anchor Rv0175
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000191|Rv0178|
MEDQQSASGDLTQKSVANGESTDTASAATEGHRGEIDAAGEPDERGAAVADSQADEDDSAATAARGGKTRARRSRGRRLAITVGVAAALFVGSAAFAGATVEPYLSERAVVATKLMVARTAANAITTLWTYTPENMDTLADRAANYLSGDFAAQYRRFVDQIAAANKQAKITNDTEVTGAAVESLSGRDAVAIVYTNTTTTSPVTKNIPALKYLSYRLFMKRYDARWLVTRMTTITSLDLTPQV