Rv0180c Family assigned · medium auto-curated

H37Rv Rv0180c · MTBC0 mtbc0_000193 · 452 aa · 211240–212598 (-) · RefSeq NP_214694.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)transmembrane protein
MTBC0 PGAP re-annotationYhgE/Pip domain-containing protein
Revised (this work)YhgE/Pip domain-containing protein. Pfam: ABC2_membrane_3 (PF12698.14), DUF3533 (PF12051.15).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O07424 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable conserved transmembrane protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionProtein of unknown function (DUF3533)
Orthologous groupCOG1511

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains) pseudogene candidate

pN/pS 0.367 · purifying
Polymorphic sites (≥ 0.1% of strains) 14 synonymous, 13 missense, 1 nonsense, 1 frameshift
Disruption 2 distinct premature-stop/frameshift site(s); most common in 5.05% of strains (7330) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
ABC2_membrane_3PF12698.14 3.9e-1734–407 ABC-2 family transporter protein
DUF3533PF12051.15 1.5e-2436–381 Protein of unknown function (DUF3533)

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3434c (transmembrane protein), high confidence from genomic context alone (score 707 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0181c hyp hypothetical protein 968 848 ctx neighborhood:846 textmining:803
Rv1006 hyp hypothetical protein 741 741 ctx cooccurence:741
Rv1435c hyp hypothetical protein 707 708 ctx cooccurence:706
Rv3434c transmembrane protein 706 707 ctx cooccurence:705
Rv2598 hyp hypothetical protein 694 695 ctx cooccurence:690
Rv1868 hyp hypothetical protein 692 692 ctx cooccurence:692
Rv0090 membrane protein 686 687 ctx cooccurence:685
Rv0182c sigG ECF RNA polymerase sigma factor SigG 740 674 ctx neighborhood:662
Rv0185 hyp hypothetical protein 662 663 ctx neighborhood:418 cooccurence:439
Rv1748 hyp hypothetical protein 661 661 ctx cooccurence:659
Rv0179c lprO lipoprotein LprO 764 641 ctx neighborhood:639
Rv0184 hyp hypothetical protein 628 628 ctx neighborhood:423
Rv3166c hyp hypothetical protein 627 627 ctx cooccurence:627
Rv2937 drrB daunorubicin ABC transporter permease DrrB 618 618 ctx cooccurence:543
Rv2114 hyp hypothetical protein 611 611 ctx cooccurence:610

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: transmembrane protein
  • MTBC0 PGAP product: YhgE/Pip domain-containing protein
  • Pfam (hmmscan --cut_ga): ABC2_membrane_3 PF12698.14 (E=4e-17), DUF3533 PF12051.15 (E=1e-24)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214694.1)
  • Domains: Pfam-A via hmmscan --cut_ga — ABC2_membrane_3 (PF12698.14), DUF3533 (PF12051.15)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1511
  • Curated reference: UniProt O07424 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 66 functional partner(s); context anchor Rv3434c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000193|Rv0180c|
MSQAQPRPAAPNPKRNVKAIRTVRFWMAPIATTLALMSALAALYLGGILNPMTNLRHFPIALVNEDAGPAGQQIVDGLVSGLDKNKFDIRVVSPDEARRLLDTAAVYGSALIPPTFSSQLRDFGASAVTPTRTDRPAITISTNPRAGTLAASIAGQTLTRALTVVNGKVGERLTAEVAAQTGGVALAGAAAAGLASPIDVKSTAYNPLPNGTGNGLSAFYYALLLLLAGFTGSIVVSTLVDSMLGYVPAEFGPVYRFAEQVNISRFRTLLVKWAVMVVLALLTSGVYLAIAHGLGMPIPLGWQVWLYGVFAIIAVGVTSSSLIAVLGSMGLLVSMLIFVILGLPSAGATVPLEAVPAFFRWLAQFEPMHQVFLGVRSLLYLNGNADAGLSQALTMTSIGLIIGLLLGGFITHLYDRSSFHRIPGAVEMAIAVEHQAQYQARQSARESSSEQP