Rv1773c Family assigned · medium auto-curated

H37Rv Rv1773c · MTBC0 mtbc0_001887 · 248 aa · 2025031–2025777 (-) · RefSeq NP_216289.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)transcriptional regulator
MTBC0 PGAP re-annotationIclR family transcriptional regulator
Revised (this work)IclR family transcriptional regulator. Pfam: HTH_IclR (PF09339.17), MarR_2 (PF12802.14), IclR_C (PF01614.25).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O06806 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable transcriptional regulatory protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category K Transcription
eggNOG descriptionTranscriptional regulator
Orthologous groupCOG1414

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.384 · purifying
Polymorphic sites (≥ 0.1% of strains) 4 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
HTH_IclRPF09339.17 3.8e-1219–68 IclR helix-turn-helix domain
MarR_2PF12802.14 4.8e-0620–72 MarR family
IclR_CPF01614.25 5.1e-4584–245 Bacterial transcriptional regulator IclR C-terminal

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0494 (HTH-type transcriptional regulator), high confidence from genomic context alone (score 869 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0494 HTH-type transcriptional regulator 875 869 ctx cooccurence:581 coexpression:693
Rv0691c mftR mycofactocin biosynthesis transcriptional regulator MftR 840 836 coexpression:836
Rv1776c transcriptional regulator 837 831 coexpression:831
Rv0377 HTH-type transcriptional regulator 836 828 coexpression:744
Rv1267c embR transcriptional regulator EmbR 834 826 coexpression:800
Rv1931c transcriptional regulator 833 825 coexpression:799
Rv1775 hyp hypothetical protein 825 825 ctx neighborhood:742
Rv3736 AraC/XylS family transcriptional regulator 828 819 coexpression:819
Rv3183 higA3 transcriptional regulator 819 813 coexpression:812
Rv3263 DNA methylase 813 813 coexpression:813
Rv1725c hyp hypothetical protein 807 808 coexpression:808
Rv1985c lysG HTH-type transcriptional regulator 816 806 coexpression:806
Rv1151c cobB NAD-dependent protein deacylase 814 806 coexpression:803
Rv1359 transcriptional regulator 805 805 coexpression:805
Rv0602c tcrA two component DNA binding transcriptional regulator TcrA 815 804 coexpression:804

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: transcriptional regulator
  • MTBC0 PGAP product: IclR family transcriptional regulator
  • Pfam (hmmscan --cut_ga): HTH_IclR PF09339.17 (E=4e-12), MarR_2 PF12802.14 (E=5e-06), IclR_C PF01614.25 (E=5e-45)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_216289.1)
  • Domains: Pfam-A via hmmscan --cut_ga — HTH_IclR (PF09339.17), MarR_2 (PF12802.14), IclR_C (PF01614.25)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1414
  • Curated reference: UniProt O06806 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 71 functional partner(s); context anchor Rv0494
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001887|Rv1773c|
MPPTEGKSTTNRDEGIQVLRRAVAALDEIAAEPGHLRLVDLCERLGLAKSTTRRLLVGLVEVGLVSVDSHGRFALGERLLGFGSVTGAHIAAAFRPTVERVARATDGETVDLSVLRGQRMWFVDQIESSYRLRAVSAVGLRFPLNGTANGKAALAALDDADAEAALCRLDPMVAEGLRREIVEIRRTGIAFDRNEHTPGISAAAIARRALGDNVIAISVPAPTARFLEKEQRIIAALRAAADSPDWTR