lpqQ Family assigned · medium auto-curated

H37Rv Rv0835 · MTBC0 mtbc0_000889 · 214 aa · 934104–934748 (+) · RefSeq NP_215350.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)lipoprotein LpqQ
MTBC0 PGAP re-annotationsensor domain-containing protein
Revised (this work)Sensor domain-containing protein. Pfam: PknH_C (PF14032.13).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt O53846 TrEMBL · unreviewed · Evidence at protein level
UniProt namePossible lipoprotein LpqQ

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
Preferred namelpqQ
eggNOG descriptionPknH-like extracellular domain
Orthologous group2B2XV

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains) pseudogene candidate

pN/pS 2.625 · diversifying/relaxed
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 8 missense, 0 nonsense, 3 frameshift
Disruption 3 distinct premature-stop/frameshift site(s); most common in 1.08% of strains (1568) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
PknH_CPF14032.13 6.3e-2137–212 PknH-like extracellular domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3198A (glutaredoxin protein), medium confidence from genomic context alone (score 545 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3198A glutaredoxin protein 545 545 ctx neighborhood:544
Rv3576 lppH lipoprotein LppH 630 65 textmining:621
Rv0556 transmembrane protein 860 47 textmining:860
Rv3705c hyp hypothetical protein 810 47 textmining:809
Rv3016 lpqA lipoprotein LpqA 806 47 textmining:805
Rv0963c hyp hypothetical protein 804 47 textmining:803
Rv1022 lpqU lipoprotein LpqU 860 46 textmining:860
Rv0403c mmpS1 membrane protein MmpS1 660 46 textmining:659
Rv1418 lprH lipoprotein LprH 870 44 textmining:870
Rv0361 membrane protein 860 44 textmining:860
Rv0178 Mce associated membrane protein 440 44 textmining:439
Rv1754c hyp hypothetical protein 628 42 textmining:628
Rv3069 crcB fluoride ion transporter CrcB 860 41 textmining:860
Rv3630 integral membrane protein 803 41 textmining:803
Rv0200 transmembrane protein 630 41 textmining:630

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: lipoprotein LpqQ
  • MTBC0 PGAP product: sensor domain-containing protein
  • Pfam (hmmscan --cut_ga): PknH_C PF14032.13 (E=6e-21)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215350.1)
  • Domains: Pfam-A via hmmscan --cut_ga — PknH_C (PF14032.13)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 2B2XV
  • Curated reference: UniProt O53846 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 17 functional partner(s); context anchor Rv3198A
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000889|Rv0835|lpqQ
MCCSTAAKSAVIVCCAAIATTACSFQATSTQPSTAPPTSRVDSLIVSIEDVRRIANYEELAAHFQTDLREPPEADTNVPGPCRVVGSSDRTFGTDWSEFRSAGYHGVTDDLRPGGPVMVETVSQAIALYPDPSTARGVFHRLESSLAECAGLHDPYFDFILDRPDASTVRIGAAGWSHVYRLKSSVFISVGVLGIEPAEPIANVILQTISDRIQ