Rv3058c Family assigned · medium auto-curated

H37Rv Rv3058c · MTBC0 mtbc0_003250 · 216 aa · 3440022–3440672 (-) · RefSeq NP_217574.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)TetR family transcriptional regulator
MTBC0 PGAP re-annotationTetR/AcrR family transcriptional regulator
Revised (this work)TetR/AcrR family transcriptional regulator. Pfam: TetR_N (PF00440.30), EthR_C (PF21313.3).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P95100 TrEMBL · unreviewed · Evidence at protein level
UniProt namePossible transcriptional regulatory protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category K Transcription
eggNOG descriptiontetR family
Orthologous groupCOG1309

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.664 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
TetR_NPF00440.30 2.4e-1226–71 Bacterial regulatory proteins, tetR family
EthR_CPF21313.3 1.2e-06102–211 Transcriptional regulator EthR, C-terminal domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3057c (short chain alcohol dehydrogenase/reductase), high confidence from genomic context alone (score 803 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3057c short chain alcohol dehydrogenase/reductase 802 803 ctx neighborhood:800
Rv3059 cyp136 cytochrome P450 Cyp136 755 754 ctx neighborhood:751
Rv0273c transcriptional regulator 717 717 ctx cooccurence:713
Rv3810 pirG cell surface protein 621 621 ctx cooccurence:615
Rv3446c hyp hypothetical protein 571 571 ctx cooccurence:569
Rv3492c Mce associated protein 510 510 ctx cooccurence:502
Rv0164 TB18.5 hyp hypothetical protein 507 508 ctx cooccurence:502
Rv1836c hyp hypothetical protein 494 494 ctx cooccurence:488
Rv3499c mce4A Mce family protein Mce4A 493 493 ctx cooccurence:488
Rv0233 nrdB ribonucleoside-diphosphate reductase subunit beta NrdB 479 479 ctx cooccurence:473
Rv0262c aac aminoglycoside 2'-N-acetyltransferase 476 476 ctx cooccurence:473
Rv0290 eccD3 ESX-3 secretion system protein EccD 475 476 ctx cooccurence:474
Rv3035 hyp hypothetical protein 448 446 ctx cooccurence:443
Rv1254 acyltransferase 429 429 ctx cooccurence:429
Rv0518 hyp hypothetical protein 425 426 ctx cooccurence:425

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: TetR family transcriptional regulator
  • MTBC0 PGAP product: TetR/AcrR family transcriptional regulator
  • Pfam (hmmscan --cut_ga): TetR_N PF00440.30 (E=2e-12), EthR_C PF21313.3 (E=1e-06)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217574.1)
  • Domains: Pfam-A via hmmscan --cut_ga — TetR_N (PF00440.30), EthR_C (PF21313.3)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1309
  • Curated reference: UniProt P95100 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 19 functional partner(s); context anchor Rv3057c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003250|Rv3058c|
MTSHAADEKQAAPPMRRRGDRHRQAILRAARELLEETPFAELSVRAISLRAGVARSGFYFYFDSKYSVLAQILAEATEELEEASQHFSARQPGESPEQFVNRMIGSVAAVYANNDPVLRACNAARQSDMEIRDILERQFQVLLRETIGVFEAEVKAGTAHPISEDLPTLVRTLAATTALMLTGDALLVGPDSDAARRVRVLEQMWLNALWGGGKAP