fusA1 Resolved · high auto-curated

H37Rv Rv0684 · MTBC0 - · 701 aa · 782485–784590 (+) · RefSeq YP_177746.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)elongation factor G
MTBC0 PGAP re-annotation
Revised (this work)Elongation factor G. Pfam: GTP_EFTU (PF00009.34), EF-G_D2 (PF22042.3), GTP_EFTU_D2 (PF03144.32), EFG_III (PF14492.13), EFG_IV (PF03764.25), EFG_C (PF00679.31).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Annotated on the H37Rv protein: this gene has no 1:1 ancestral MTBC0 anchor (PE/PPE, paralogue, IS element, or otherwise unanchored CDS).

Curated reference (UniProt)

UniProt P9WNM7 SwissProt · reviewed · Evidence at protein level
UniProt nameElongation factor G
Curated functionCatalyzes the GTP-dependent ribosomal translocation step during translation elongation. During this step, the ribosome changes from the pre-translocational (PRE) to the post-translocational (POST) state as the newly formed A-site-bound peptidyl-tRNA and P-site-bound deacylated tRNA move to the P and E sites, respectively. Catalyzes the coordinated movement of the two tRNA molecules, the mRNA and conformational changes in the ribosome (By similarity).

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category J Translation, ribosomal structure and biogenesis
Preferred namefusA
eggNOG descriptionCatalyzes the GTP-dependent ribosomal translocation step during translation elongation. During this step, the ribosome changes from the pre-translocational (PRE) to the post- translocational (POST) state as the newly formed A-site-bound peptidyl-tRNA and P-site-bound deacylated tRNA move to the P and E sites, respectively. Catalyzes the coordinated movement of the two tRNA molecules, the mRNA and conformational changes in the ribosome
Orthologous groupCOG0480
KEGG orthology K02355
Gene Ontology (10) GO:0005575, GO:0005618, GO:0005623, GO:0005886, GO:0008150, GO:0016020, GO:0030312, GO:0040007, GO:0044464, GO:0071944

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.044 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 14 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
GTP_EFTUPF00009.34 3.0e-6512–285 Elongation factor Tu GTP binding domain
EF-G_D2PF22042.3 3.9e-15313–395 Elongation factor G domain 2
GTP_EFTU_D2PF03144.32 3.3e-16328–394 Elongation factor Tu domain 2
EFG_IIIPF14492.13 2.2e-33408–482 Elongation Factor G, domain III
EFG_IVPF03764.25 3.0e-45483–604 Elongation factor G, domain IV
EFG_CPF00679.31 5.3e-30608–692 Elongation factor G C-terminus

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: rpsG (30S ribosomal protein S7), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0683 rpsG 30S ribosomal protein S7 999 1000 ctx neighborhood:784 cooccurence:722 coexpression:943 experimental:829 textmining:861
Rv0682 rpsL 30S ribosomal protein S12 999 1000 ctx neighborhood:784 cooccurence:695 coexpression:945 experimental:829 textmining:783
Rv0704 rplB 50S ribosomal protein L2 999 998 ctx cooccurence:707 coexpression:949 experimental:829 textmining:776
Rv0702 rplD 50S ribosomal protein L4 998 998 ctx cooccurence:622 coexpression:945 experimental:829
Rv0685 tuf elongation factor Tu 998 997 ctx neighborhood:728 coexpression:937 experimental:778 textmining:642
Rv0701 rplC 50S ribosomal protein L3 997 997 ctx cooccurence:627 coexpression:938 experimental:829
Rv0714 rplN 50S ribosomal protein L14 998 996 ctx cooccurence:560 coexpression:924 experimental:829 textmining:570
Rv0716 rplE 50S ribosomal protein L5 997 996 ctx cooccurence:547 coexpression:912 experimental:829 textmining:440
Rv0719 rplF 50S ribosomal protein L6 997 996 coexpression:951 experimental:829 textmining:406
Rv0700 rpsJ 30S ribosomal protein S10 997 996 ctx cooccurence:432 coexpression:944 experimental:829
Rv0705 rpsS 30S ribosomal protein S19 996 996 ctx cooccurence:463 coexpression:931 experimental:829
Rv0718 rpsH 30S ribosomal protein S8 996 996 coexpression:943 experimental:829
Rv3460c rpsM 30S ribosomal protein S13 996 995 ctx cooccurence:555 coexpression:931 experimental:773
Rv3458c rpsD 30S ribosomal protein S4 996 995 coexpression:930 experimental:829
Rv0715 rplX 50S ribosomal protein L24 995 995 coexpression:943 experimental:829

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Annotation from H37Rv (no MTBC0 1:1 anchor; H37Rv protein used): elongation factor G
  • Pfam (hmmscan --cut_ga): GTP_EFTU PF00009.34 (E=3e-65), EF-G_D2 PF22042.3 (E=4e-15), GTP_EFTU_D2 PF03144.32 (E=3e-16), EFG_III PF14492.13 (E=2e-33), EFG_IV PF03764.25 (E=3e-45), EFG_C PF00679.31 (E=5e-30)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq YP_177746.1)
  • Domains: Pfam-A via hmmscan --cut_ga — GTP_EFTU (PF00009.34), EF-G_D2 (PF22042.3), GTP_EFTU_D2 (PF03144.32), EFG_III (PF14492.13), EFG_IV (PF03764.25), EFG_C (PF00679.31)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0480
  • Curated reference: UniProt P9WNM7 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 262 functional partner(s); context anchor rpsG
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>H37Rv|Rv0684|fusA1
MAQKDVLTDLSRVRNFGIMAHIDAGKTTTTERILYYTGINYKIGEVHDGAATMDWMEQEQERGITITSAATTTFWKDNQLNIIDTPGHVDFTVEVERNLRVLDGAVAVFDGKEGVEPQSEQVWRQADKYDVPRICFVNKMDKIGADFYFSVRTMGERLGANAVPIQLPVGAEADFEGVVDLVEMNAKVWRGETKLGETYDTVEIPADLAEQAEEYRTKLLEVVAESDEHLLEKYLGGEELTVDEIKGAIRKLTIASEIYPVLCGSAFKNKGVQPMLDAVVDYLPSPLDVPPAIGHAPAKEDEEVVRKATTDEPFAALAFKIATHPFFGKLTYIRVYSGTVESGSQVINATKGKKERLGKLFQMHSNKENPVDRASAGHIYAVIGLKDTTTGDTLSDPNQQIVLESMTFPDPVIEVAIEPKTKSDQEKLSLSIQKLAEEDPTFKVHLDSETGQTVIGGMGELHLDILVDRMRREFKVEANVGKPQVAYKETIKRLVQNVEYTHKKQTGGSGQFAKVIINLEPFTGEEGATYEFESKVTGGRIPREYIPSVDAGAQDAMQYGVLAGYPLVNLKVTLLDGAYHEVDSSEMAFKIAGSQVLKKAAALAQPVILEPIMAVEVTTPEDYMGDVIGDLNSRRGQIQAMEERAGARVVRAHVPLSEMFGYVGDLRSKTQGRANYSMVFDSYSEVPANVSKEIIAKATGE