sdaA Resolved · high auto-curated

H37Rv Rv0069c · MTBC0 mtbc0_000076 · 461 aa · 76382–77767 (-) · RefSeq NP_214583.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)L-serine dehydratase
MTBC0 PGAP re-annotationL-serine ammonia-lyase
Revised (this work)L-serine ammonia-lyase. Pfam: SDH_beta (PF03315.22), SDH_alpha (PF03313.22).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WGT5 SwissProt · reviewed · Evidence at protein level
UniProt nameL-serine dehydratase
EC (curated) EC 4.3.1.17

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category E Amino acid transport and metabolism
Preferred namesdaA
eggNOG descriptionL-serine dehydratase
Orthologous groupCOG1760
EC number EC 4.3.1.17
KEGG orthology K01752
KEGG pathways map00260, map00270, map01100, map01110, map01130, map01200, map01230
Gene Ontology (12) GO:0003674, GO:0003824, GO:0003941, GO:0005575, GO:0005623, GO:0005886, GO:0016020, GO:0016829, GO:0016840, GO:0016841, GO:0044464, GO:0071944

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 3.142 · diversifying/relaxed
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 9 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
SDH_betaPF03315.22 4.4e-554–156 Serine dehydratase beta chain
SDH_alphaPF03313.22 1.5e-90187–451 Serine dehydratase alpha chain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: glyA2 (serine hydroxymethyltransferase), high confidence from genomic context alone (score 997 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0070c glyA2 serine hydroxymethyltransferase 998 997 ctx neighborhood:801 coexpression:839 database:900 textmining:518
Rv1093 glyA1 serine hydroxymethyltransferase 964 925 database:900 textmining:547
Rv1559 ilvA threonine dehydratase IlvA 960 922 database:900 textmining:518
Rv1077 cbs cystathionine beta-synthase 926 905 database:900
Rv1613 trpA tryptophan synthase subunit alpha 908 905 database:900
Rv3042c serB2 phosphoserine phosphatase SerB 973 904 database:900 textmining:737
Rv0436c pssA CDP-diacylglycerol--serine O-phosphatidyltransferase 901 902 database:900
Rv2335 cysE serine acetyltransferase 956 901 database:900 textmining:579
Rv1612 trpB tryptophan synthase subunit beta 904 901 database:900
Rv1296 thrB homoserine kinase 860 843 database:800
Rv1294 thrA homoserine dehydrogenase 846 839 database:800
Rv3341 metA homoserine O-acetyltransferase 800 801 database:800
Rv1832 gcvB glycine dehydrogenase 872 668 coexpression:403 textmining:631
Rv3002c ilvN acetolactate synthase small subunit 651 631 database:500
Rv1295 thrC threonine synthase 636 600 database:500

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: L-serine dehydratase
  • MTBC0 PGAP product: L-serine ammonia-lyase
  • Pfam (hmmscan --cut_ga): SDH_beta PF03315.22 (E=4e-55), SDH_alpha PF03313.22 (E=2e-90)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214583.1)
  • Domains: Pfam-A via hmmscan --cut_ga — SDH_beta (PF03315.22), SDH_alpha (PF03313.22)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1760
  • Curated reference: UniProt P9WGT5 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 38 functional partner(s); context anchor glyA2
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000076|Rv0069c|sdaA
MTISVFDLFTIGIGPSSSHTVGPMRAANQFVVALRRRGHLDDLEAMRVDLFGSLAATGAGHGTMSAILLGLEGCQPETITTEHKERRLAEIAASGVTRIGGVIPVPLTERDIDLHPDIVLPTHPNGMTFTAAGPHGRVLATETYFSVGGGFIVTEQTSGNSGQHPCSVALPYVSAQELLDICDRLDVSISEAALRNETCCRTENEVRAALLHLRDVMVECEQRSIAREGLLPGGLRVRRRAKVWYDRLNAEDPTRKPEFAEDWVNLVALAVNEENASGGRVVTAPTNGAAGIVPAVLHYAIHYTSAGAGDPDDVTVRFLLTAGAIGSLFKERASISGAEVGCQGEVGSAAAMAAAGLAEILGGTPRQVENAAEIAMEHSLGLTCDPIAGLVQIPCIERNAISAGKAINAARMALRGDGIHRVTLDQVIDTMRATGADMHTKYKETSAGGLAINVAVNIVEC