yidD Resolved · high auto-curated

H37Rv Rv3922c · MTBC0 mtbc0_004156 · 120 aa · 4434304–4434666 (-) · RefSeq NP_218439.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)membrane protein insertion efficiency factor
MTBC0 PGAP re-annotationmembrane protein insertion efficiency factor YidD
Revised (this work)Membrane protein insertion efficiency factor YidD. Pfam: YidD (PF01809.24).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WFL9 SwissProt · reviewed · Inferred from homology
UniProt namePutative membrane protein insertion efficiency factor
Curated functionCould be involved in insertion of integral membrane proteins into the membrane.

UniProt still lists this protein as Putative membrane protein insertion efficiency factor; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
Preferred nameytjA
eggNOG descriptionCould be involved in insertion of integral membrane proteins into the membrane
Orthologous groupCOG0759
KEGG orthology K08998

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.663 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 4 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
YidDPF01809.24 5.1e-2822–86 Putative membrane protein insertion efficiency factor

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: rnpA (ribonuclease P protein component), high confidence from genomic context alone (score 996 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3923c rnpA ribonuclease P protein component 999 996 ctx neighborhood:881 coexpression:968 textmining:885
Rv3921c yidC membrane protein insertase YidC 998 988 ctx neighborhood:881 coexpression:881 textmining:845
Rv3920c hyp hypothetical protein 955 955 ctx neighborhood:792 coexpression:794
Rv3924c rpmH 50S ribosomal protein L34 966 876 ctx neighborhood:816 textmining:739
Rv1297 rho transcription termination factor Rho 857 857 coexpression:825
Rv0704 rplB 50S ribosomal protein L2 782 783 coexpression:783
Rv0702 rplD 50S ribosomal protein L4 774 774 coexpression:774
Rv0638 secE1 preprotein translocase SecE 746 746 coexpression:746
Rv3834c serS serine--tRNA ligase 728 728 coexpression:719
Rv3918c parA chromosome partitioning protein ParA 697 697 ctx neighborhood:688
Rv2966c rsmD methyltransferase 700 695 coexpression:685
Rv3917c parB chromosome partitioning protein ParB 662 663 ctx neighborhood:642
Rv0684 fusA1 elongation factor G 617 617 coexpression:615
Rv3919c gid 16S rRNA (guanine(527)-N(7))-methyltransferase RsmG 605 606 ctx neighborhood:501
Rv0979A rpmF 50S ribosomal protein L32 557 537 coexpression:537

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: membrane protein insertion efficiency factor
  • MTBC0 PGAP product: membrane protein insertion efficiency factor YidD
  • Pfam (hmmscan --cut_ga): YidD PF01809.24 (E=5e-28)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218439.1)
  • Domains: Pfam-A via hmmscan --cut_ga — YidD (PF01809.24)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0759
  • Curated reference: UniProt P9WFL9 (SwissProt, reviewed; Inferred from homology)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 31 functional partner(s); context anchor rnpA
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_004156|Rv3922c|yidD
MSLSRQSCGRVVRVTGRASARGLIFVIQVYRHMLSPLRPASCRFVPTCSQYAVDALTEYGLLRGSWLTMIRLAKCGPWHRGGWDPIPEGLTTGRSCQTDVDGANDDWNPASKRGERESFV