Rv3492c Resolved · medium auto-curated

H37Rv Rv3492c · MTBC0 mtbc0_003707 · 160 aa · 3935047–3935529 (-) · RefSeq NP_218009.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)Mce associated protein
MTBC0 PGAP re-annotationMce associated protein
Revised (this work)Mce associated protein.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt I6YGA5 TrEMBL · unreviewed · Evidence at protein level
UniProt nameConserved hypothetical Mce associated protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionProtein conserved in bacteria
Orthologous groupCOG3184
KEGG orthology K18481
KEGG modules M00670

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.216 · purifying
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: mce4D (Mce family protein Mce4D), high confidence from genomic context alone (score 990 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3496c mce4D Mce family protein Mce4D 990 990 ctx neighborhood:879 cooccurence:723 coexpression:734
Rv3495c lprN Mce family lipoprotein LprN 978 978 ctx neighborhood:879 cooccurence:755
Rv3494c mce4F Mce family protein Mce4 974 974 ctx neighborhood:881 cooccurence:723
Rv3493c Mce associated protein 973 973 ctx neighborhood:882 coexpression:780
Rv3499c mce4A Mce family protein Mce4A 975 971 ctx neighborhood:879 cooccurence:762
Rv3497c mce4C Mce family protein Mce4C 989 970 ctx neighborhood:879 cooccurence:708 textmining:657
Rv3498c mce4B Mce family protein Mce4B 958 959 ctx neighborhood:879 cooccurence:650
Rv3500c yrbE4B integral membrane protein 917 917 ctx neighborhood:856 cooccurence:440
Rv3501c yrbE4A integral membrane protein 883 883 ctx neighborhood:813
Rv3529c hyp hypothetical protein 809 801 experimental:417 database:540
Rv1363c membrane protein 770 771 ctx cooccurence:764
Rv0169 mce1A Mce family protein Mce1A 796 756 ctx cooccurence:735
Rv1972 Mce associated membrane protein 755 755 ctx cooccurence:752
Rv2390c hyp hypothetical protein 753 754 ctx cooccurence:752
Rv0172 mce1D Mce family protein Mce1D 853 751 ctx cooccurence:683 textmining:435

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: Mce associated protein
  • MTBC0 PGAP product: Mce associated protein
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_218009.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG3184
  • Curated reference: UniProt I6YGA5 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 314 functional partner(s); context anchor mce4D
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003707|Rv3492c|
MRRLISVAYALMVATIVGLSAAGGWFYWDRVQTGGEASARALLPKLAMQEIPQVFGYDYQTVERSLTAVYPLLTPDYRQEFQKSANAQIIPEAKKREVVVQANVVGVGVMDAKRDCASVMVYLNRTVTDKTRQPLYDGSRLRVDFQRIDGKWLIAYITPI