Rv1363c Still unknown · low auto-curated

H37Rv Rv1363c · MTBC0 mtbc0_001465 · 261 aa · 1545309–1546094 (-) · RefSeq NP_215879.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)membrane protein
MTBC0 PGAP re-annotationhypothetical protein
Revised (this work)Conserved hypothetical protein; no recognised domain. Function unknown.

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WLZ9 SwissProt · reviewed · Evidence at protein level
UniProt nameUncharacterized protein Rv1363c

UniProt still lists this protein as Uncharacterized protein Rv1363c; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

Orthologous group2E5BU
KEGG orthology K18481
KEGG modules M00670

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.643 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 3 synonymous, 6 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

No Pfam-A domain above the gathering threshold (or not yet scanned).

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv1362c (membrane protein), high confidence from genomic context alone (score 987 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1362c membrane protein 986 987 ctx neighborhood:800 cooccurence:773 coexpression:732
Rv3492c Mce associated protein 770 771 ctx cooccurence:764
Rv1973 Mce associated membrane protein 760 760 ctx cooccurence:726
Rv0172 mce1D Mce family protein Mce1D 758 758 ctx cooccurence:747
Rv0169 mce1A Mce family protein Mce1A 751 751 ctx cooccurence:740
Rv0594 mce2F Mce family protein Mce2F 750 751 ctx cooccurence:740
Rv0174 mce1F Mce family protein Mce1F 748 749 ctx cooccurence:738
Rv3499c mce4A Mce family protein Mce4A 745 745 ctx cooccurence:733
Rv3825c pks2 phthioceranic/hydroxyphthioceranic acid synthase 764 736 experimental:470 database:508
Rv2048c pks12 polyketide synthase 763 735 experimental:470 database:508
Rv1527c pks5 polyketide synthase 763 735 experimental:470 database:508
Rv2940c mas multifunctional mycocerosic acid synthase 763 735 experimental:470 database:508
Rv2933 ppsC phthiocerol synthesis polyketide synthase type I PpsC 762 735 experimental:470 database:508
Rv2267c stf3 hyp hypothetical protein 746 735 experimental:417 database:540
Rv3446c hyp hypothetical protein 743 733

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: membrane protein
  • MTBC0 PGAP product: hypothetical protein
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215879.1)
  • Domains: Pfam-A via hmmscan --cut_ga — none above threshold
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG 2E5BU
  • Curated reference: UniProt P9WLZ9 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 303 functional partner(s); context anchor Rv1362c
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001465|Rv1363c|
MAETTEPPSDAGTSQADAMALAAEAEAAEAEALAAAARARARAARLKREALAMAPAEDENVPEEYADWEDAEDYDDYDDYEAADQEAARSASWRRRLRVRLPRLSTIAMAAAVVIICGFTGLSGYIVWQHHEATERQQRAAAFAAGAKQGVINMTSLDFNKAKEDVARVIDSSTGEFRDDFQQRAADFTKVVEQSKVVTEGTVNATAVESMNEHSAVVLVAATSRVTNSAGAKDEPRAWRLKVTVTEEGGQYKMSKVEFVP