Rv1376 Family assigned · medium auto-curated

H37Rv Rv1376 · MTBC0 mtbc0_001477 · 497 aa · 1558492–1559985 (+) · RefSeq NP_215892.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationTfuA-like protein
Revised (this work)TfuA-like protein. Pfam: TfuA (PF07812.19).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P71804 TrEMBL · unreviewed · Predicted
UniProt nameTfuA-like core domain-containing protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category S Function unknown
eggNOG descriptionTfuA-like protein
Orthologous groupCOG3482

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.855 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 5 synonymous, 12 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
TfuAPF07812.19 5.9e-4353–172 TfuA-like protein

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: PE_PGRS28 (PE-PGRS family protein PE_PGRS28), medium confidence from genomic context alone (score 688 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv1375 hyp hypothetical protein 988 986 ctx neighborhood:882 cooccurence:774 coexpression:533
Rv1452c PE_PGRS28 PE-PGRS family protein PE_PGRS28 688 688 ctx cooccurence:688
Rv3347c PPE55 PPE family protein PPE55 675 676 ctx cooccurence:674
Rv2490c PE_PGRS43 PE-PGRS family protein PE_PGRS43 670 670 ctx cooccurence:670
Rv3350c PPE56 PPE family protein PPE56 669 669 ctx cooccurence:668
Rv0355c PPE8 PPE family protein PPE8 668 668 ctx cooccurence:667
Rv2337c hyp hypothetical protein 648 649 ctx cooccurence:636
Rv2209 integral membrane protein 648 648 ctx cooccurence:646
Rv3343c PPE54 PPE family protein PPE54 641 641 ctx cooccurence:639
Rv1917c PPE34 PPE family protein PPE34 637 637 ctx cooccurence:635
Rv1651c PE_PGRS30 PE-PGRS family protein PE_PGRS30 634 634 ctx cooccurence:634
Rv1004c membrane protein 628 628 ctx cooccurence:628
Rv2819c csm5 CRISPR type III-associated RAMP protein Csm5 626 626 ctx cooccurence:621
Rv0304c PPE5 PPE family protein PPE5 612 612 ctx cooccurence:612
Rv2293c hyp hypothetical protein 588 589 ctx cooccurence:586

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: TfuA-like protein
  • Pfam (hmmscan --cut_ga): TfuA PF07812.19 (E=6e-43)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215892.1)
  • Domains: Pfam-A via hmmscan --cut_ga — TfuA (PF07812.19)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG3482
  • Curated reference: UniProt P71804 (TrEMBL, unreviewed; Predicted)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 65 functional partner(s); context anchor PE_PGRS28
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001477|Rv1376|
MTACGRIVVTAGPTISAADIRSVVPDAEVAPPIAFGQALSYDLRSGDTLLIVDGLFFQQPSVRHKELLTLMADGVRVVGSSSMGALRAAELHPFGMEGYGWVFESYRDGVLEADDEVGVVHGDADDGYPVFVDALVNMRHTLARAVATGVVCSELAERIIETARATPFTMRTWARLLSEVGAPDQRGLAAQLRSLRVDVKHADALLALRQLGQRPRVEPLRPGPPPTVWSRRWRQRWAPPTSVAASADHGESFVDVTDLEVLSFLSVSSVDYWAYRPALQQVAAWYWTLKHPEQSGSVGERAARAVAEVASEGYGRALEFIAYRYALATGIIDETGFPEAVAAHWLTTEERHGLGNDPISISARVITRTLFVVRLLPAIDHFLDLLRKDSRLPRWRAMAAHALCKRDDLARQKPHLNLGRPDPTQLKRLFGARWGTQVNRIELARRGLMTEDAFYAAATPFAVAAVDDQLPRIEVGTLGPAPLSADVPERHFDFGSV