Rv3271c Resolved · high auto-curated

H37Rv Rv3271c · MTBC0 mtbc0_003479 · 222 aa · 3674822–3675490 (-) · RefSeq NP_217788.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)integral membrane protein
MTBC0 PGAP re-annotationcation transporter
Revised (this work)Cation transporter. Pfam: Cation_efflux (PF01545.27).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P96876 TrEMBL · unreviewed · Evidence at protein level
UniProt nameProbable conserved integral membrane protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category P Inorganic ion transport and metabolism
eggNOG descriptionCation efflux family
Orthologous groupCOG0053

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.368 · purifying
Polymorphic sites (≥ 0.1% of strains) 1 synonymous, 1 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
Cation_effluxPF01545.27 8.0e-0637–213 Cation efflux transmembrane domain

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv3273 (transmembrane carbonic anhydrase), medium confidence from genomic context alone (score 603 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3272 hyp hypothetical protein 782 782 ctx neighborhood:781
Rv1240 mdh malate dehydrogenase 682 683 coexpression:672
Rv3273 transmembrane carbonic anhydrase 619 603 ctx neighborhood:597
Rv0648 alpha-mannosidase 510 510 coexpression:411
Rv2159c hyp hypothetical protein 427 427
Rv1712 cmk cytidylate kinase 423 424 coexpression:405
Rv1994c cmtR HTH-type transcriptional regulator CmtR 419 391
Rv2683 hyp hypothetical protein 410 384
Rv3483c hyp hypothetical protein 671 147 textmining:630
Rv1275 lprC lipoprotein LprC 630 85 textmining:613
Rv2182c 1-acylglycerol-3-phosphate O-acyltransferase 461 84 textmining:436
Rv1457c antibiotic ABC transporter permease 476 77 textmining:457
Rv0625c transmembrane protein 669 73 textmining:658
Rv1337 integral membrane protein 638 63 textmining:630
Rv0227c membrane protein 552 55 textmining:546

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: integral membrane protein
  • MTBC0 PGAP product: cation transporter
  • Pfam (hmmscan --cut_ga): Cation_efflux PF01545.27 (E=8e-06)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217788.1)
  • Domains: Pfam-A via hmmscan --cut_ga — Cation_efflux (PF01545.27)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0053
  • Curated reference: UniProt P96876 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 23 functional partner(s); context anchor Rv3273
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003479|Rv3271c|
METTTEHRDESTLDSPVSVAREAEWQRNVRWARWLAWVSLAVLLTEGAVGLWQGIAVGSVALTGWALGGGSEGLASAMVLWRFTGDRTWSATAEHRAQRGVAVSFWLTAPYLVAESIRHLAGEHRAETSVIGIGLTAIALLLMPVLGWANHRVGERLGSGATAGEGTQNYLCAAQAAAVLLGLAITAVWSNGWWIDPAIGLAIAGIAVWQGIRTWRGHGCGC