lcp1 Resolved · high auto-curated

H37Rv Rv3267 · MTBC0 mtbc0_003475 · 498 aa · 3670028–3671524 (+) · RefSeq NP_217784.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)hypothetical protein
MTBC0 PGAP re-annotationphosphotransferase Lcp1
Revised (this work)Phosphotransferase Lcp1. Pfam: LytR_cpsA_psr (PF03816.20), LytR_C (PF13399.12).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P96872 TrEMBL · unreviewed · Evidence at protein level
UniProt nameConserved protein

UniProt still lists this protein as Conserved protein; the revised annotation above is ahead of the current UniProt record.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category K Transcription
Preferred namelytR2
eggNOG descriptioncell envelope-related transcriptional attenuator
Orthologous groupCOG1316

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS n/a
Polymorphic sites (≥ 0.1% of strains) 0 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
LytR_cpsA_psrPF03816.20 1.6e-4295–263 LytR_cpsA_psr family
LytR_CPF13399.12 2.6e-14365–449 LytR cell envelope-related transcriptional attenuator

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: manB (D-alpha-D-mannose-1-phosphate guanylyltransferase ManB), high confidence from genomic context alone (score 847 excluding text-mining). This association is the citable seed of a function hypothesis for this hypothetical protein.

PartnerProductScoreNo text-miningChannels (≥400)
Rv3268 hyp hypothetical protein 951 951 ctx neighborhood:825 coexpression:734
Rv3264c manB D-alpha-D-mannose-1-phosphate guanylyltransferase ManB 847 847 ctx neighborhood:783
Rv3265c wbbL1 N-acetylglucosaminyl-diphospho-decaprenol L-rhamnosyltransferase 919 831 ctx neighborhood:783 textmining:545
Rv3266c rmlD dTDP-4-dehydrorhamnose reductase 931 787 ctx neighborhood:784 textmining:693
Rv1697 steA hyp hypothetical protein 766 766 coexpression:732
Rv3270 ctpC manganese/zinc-exporting P-type ATPase 716 717 ctx neighborhood:715
Rv3269 hyp hypothetical protein 688 689 ctx neighborhood:687
Rv3311 hyp hypothetical protein 649 649 ctx cooccurence:644
Rv3212 hyp hypothetical protein 602 603 ctx cooccurence:597
Rv0996 transmembrane protein 599 599 ctx cooccurence:512
Rv3438 hyp hypothetical protein 581 581 ctx cooccurence:581
Rv3794 embA arabinosyltransferase A 589 539 ctx cooccurence:533
Rv3795 embB arabinosyltransferase B 551 521 ctx cooccurence:517
Rv3244c lpqB lipoprotein LpqB 520 521 ctx cooccurence:492
Rv3793 embC arabinosyltransferase C 530 519 ctx cooccurence:512

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: hypothetical protein
  • MTBC0 PGAP product: phosphotransferase Lcp1
  • Pfam (hmmscan --cut_ga): LytR_cpsA_psr PF03816.20 (E=2e-42), LytR_C PF13399.12 (E=3e-14)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217784.1)
  • Domains: Pfam-A via hmmscan --cut_ga — LytR_cpsA_psr (PF03816.20), LytR_C (PF13399.12)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG1316
  • Curated reference: UniProt P96872 (TrEMBL, unreviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 54 functional partner(s); context anchor manB
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003475|Rv3267|lcp1
MMSAQRVVRTVRTARAISTALAVAIVLGTGVAWSSVRSFEDGIFHMSAPSLGHGGDDGAIDILLVGLDSRTDAHGNPLSAEELATLHAGDEEATNTDTIILIRVPNNGKSATAISIPRDSYVAAPGLGKTKINGVYGQTRETKRAGLVQAGASPTEAAAAGTEAGREALIKTVADLTGVTVDHYAEIGLLGFALIADALGGVDVCLKEPVYEPLSGADFPAGRQKLNGPQALSFVRQRHDLPRGDLDRVVRQQAVMAALAHRVISGQTLSSPATLKRLEQAVQRSVVLSSGWDIMDFVRQLQKLAGGNVAFATIPVLDGAGWSDDGMQSVVRVDPRQVQDWVVGLLHEQDQGKTDELAYTPAKTTANVVNDTDINGLAAAVSKVLSSKGFTTGSVGNNDGDHVPGSQVRAAKADDLGAQQVAKELGGLPVVADASIAPGSVRVVLANDYSGPGSGLGGSDPNGVVSPARAFNLGSADDTTPPPSPILTAGSDAPECIN