ftsX Resolved · high auto-curated

H37Rv Rv3101c · MTBC0 mtbc0_003297 · 297 aa · 3491150–3492043 (-) · RefSeq NP_217617.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)cell division protein FtsX
MTBC0 PGAP re-annotationpermease-like cell division protein FtsX
Revised (this work)Permease-like cell division protein FtsX. Pfam: FtsX_ECD (PF18075.9), FtsX (PF02687.28).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WG19 SwissProt · reviewed · Evidence at protein level
UniProt nameCell division protein FtsX
Curated functionPart of the ABC transporter FtsEX involved in cellular division.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category D Cell cycle control, cell division, chromosome partitioning
Preferred nameftsX
eggNOG descriptionPart of the ABC transporter FtsEX involved in cellular division
Orthologous groupCOG2177
KEGG orthology K09811
KEGG pathways map02010
KEGG modules M00256
Gene Ontology (19) GO:0005575, GO:0005576, GO:0005618, GO:0005623, GO:0005886, GO:0005887, GO:0008150, GO:0009987, GO:0016020, GO:0016021, GO:0030312, GO:0031224 +7 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.332 · purifying
Polymorphic sites (≥ 0.1% of strains) 2 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
FtsX_ECDPF18075.9 2.6e-2156–166 FtsX extracellular domain
FtsXPF02687.28 1.2e-14175–293 FtsX-like permease C-terminal

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: ftsE (cell division ATP-binding protein FtsE), high confidence from genomic context alone (score 997 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv3102c ftsE cell division ATP-binding protein FtsE 999 997 ctx neighborhood:882 cooccurence:764 coexpression:892 textmining:974
Rv3100c smpB SsrA-binding protein 960 896 ctx neighborhood:882 textmining:630
Rv3099c hyp hypothetical protein 832 832 ctx neighborhood:830
Rv3105c prfB peptide chain release factor PrfB 778 709 ctx neighborhood:706
Rv3104c transmembrane protein 706 706 ctx neighborhood:702
Rv3103c hyp hypothetical protein 690 690 ctx neighborhood:682
Rv1631 coaE dephospho-CoA kinase CoaE 533 533 coexpression:439
Rv3106 fprA NADPH-ferredoxin reductase FprA 526 526 ctx neighborhood:525
Rv0950c hyp hypothetical protein 490 453
Rv1477 ripA peptidoglycan endopeptidase RipA 610 449
Rv2553c mltG membrane protein 445 421
Rv1239c corA magnesium and cobalt transport transmembrane protein CorA 421 421 coexpression:402
Rv3915 cwlM peptidoglycan hydrolase 578 407
Rv1709 scpA segregation and condensation protein ScpA 402 403
Rv2145c wag31 cell wall synthesis protein Wag31 494 390

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: cell division protein FtsX
  • MTBC0 PGAP product: permease-like cell division protein FtsX
  • Pfam (hmmscan --cut_ga): FtsX_ECD PF18075.9 (E=3e-21), FtsX PF02687.28 (E=1e-14)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_217617.1)
  • Domains: Pfam-A via hmmscan --cut_ga — FtsX_ECD (PF18075.9), FtsX (PF02687.28)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2177
  • Curated reference: UniProt P9WG19 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 31 functional partner(s); context anchor ftsE
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_003297|Rv3101c|ftsX
MRFGFLLNEVLTGFRRNVTMTIAMILTTAISVGLFGGGMLVVRLADSSRAIYLDRVESQVFLTEDVSANDSSCDTTACKALREKIETRSDVKAVRFLNRQQAYDDAIRKFPQFKDVAGKDSFPASFIVKLENPEQHKDFDTAMKGQPGVLDVLNQKELIDRLFAVLDGLSNAAFAVALVQAIGAILLIANMVQVAAYTRRTEIGIMRLVGASRWYTQLPFLVEAMLAATMGVGIAVAGLMVVRALFLENALNQFYQANLIAKVDYADILFITPWLLLLGVAMSGLTAYLTLRLYVRR