oppA Family assigned · medium auto-curated

H37Rv Rv1280c · MTBC0 mtbc0_001370 · 591 aa · 1440648–1442423 (-) · RefSeq NP_215796.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)oligopeptide ABC transporter substrate-binding lipoprotein OppA
MTBC0 PGAP re-annotationoligopeptide ABC transporter substrate-binding protein OppA
Revised (this work)Oligopeptide ABC transporter substrate-binding protein OppA. Pfam: SBP_bac_5 (PF00496.28).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WGU5 SwissProt · reviewed · Evidence at protein level
UniProt nameOligopeptide-binding protein OppA
Curated functionPart of the ABC transporter complex OppABCD involved in the uptake of oligopeptides. Peptide-binding protein that shows broad specificity but a moderate preference for hydrophobic oligopeptides and those that are 6-16 amino acids long. Can bind the tripeptide glutathione (GSH) and the nonapeptide bradykinin. Glutathione (GSH) uptake by mycobacteria through the OppABCD system contributes to the depletion of the GSH pool in infected macrophages, which impairs the ability of the macrophage to detoxify methylglyoxal (MG) and contributes to enhanced production of inflammatory cytokines. May also pr.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category E Amino acid transport and metabolism
Preferred nameoppA
eggNOG descriptionextracellular solute-binding
Orthologous groupCOG0747
KEGG orthology K02035
KEGG pathways map02024
KEGG modules M00239
Gene Ontology (45) GO:0003674, GO:0005215, GO:0005488, GO:0005575, GO:0005623, GO:0005886, GO:0005887, GO:0006810, GO:0008104, GO:0008150, GO:0015031, GO:0015833 +33 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.575 · relaxed/neutral
Polymorphic sites (≥ 0.1% of strains) 7 synonymous, 12 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
SBP_bac_5PF00496.28 2.3e-52152–511 Bacterial extracellular solute-binding proteins, family 5 Middle

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: oppB (oligopeptide ABC transporter permease OppB), high confidence from genomic context alone (score 998 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1283c oppB oligopeptide ABC transporter permease OppB 999 998 ctx neighborhood:881 cooccurence:693 coexpression:459 database:900 textmining:914
Rv1282c oppC oligopeptide ABC transporter permease OppC 998 998 ctx neighborhood:881 cooccurence:651 coexpression:428 database:900 textmining:405
Rv1281c oppD oligopeptide ABC transporter ATP-binding protein OppD 993 993 ctx neighborhood:802 cooccurence:617 database:900
Rv3663c dppD dipeptide ABC transporter ATP-binding protein DppD 989 934 database:900 textmining:851
Rv2585c lipoprotein 913 913 database:900
Rv1284 canA beta-carbonic anhydrase 756 756 ctx neighborhood:756
Rv2455c korA 2-oxoglutarate oxidoreductase subunit KorA 669 669 coexpression:647
Rv3664c dppC dipeptide ABC transporter permease DppC 839 633 coexpression:429 textmining:580
Rv3665c dppB dipeptide ABC transporter permease DppB 793 627 coexpression:459 textmining:469
Rv2326c ABC transporter ATP-binding protein 567 540
Rv1286 cysC adenylyl-sulfate kinase 526 526
Rv2862c hyp hypothetical protein 482 482 ctx cooccurence:482
Rv3666c dppA dipeptide ABC transporter substrate-binding lipoprotein DppA 742 445 textmining:555
Rv0320 hyp hypothetical protein 442 443 ctx cooccurence:435
Rv2397c cysA1 sulfate ABC transporter ATP-binding protein CysA 442 443

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: oligopeptide ABC transporter substrate-binding lipoprotein OppA
  • MTBC0 PGAP product: oligopeptide ABC transporter substrate-binding protein OppA
  • Pfam (hmmscan --cut_ga): SBP_bac_5 PF00496.28 (E=2e-52)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215796.1)
  • Domains: Pfam-A via hmmscan --cut_ga — SBP_bac_5 (PF00496.28)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0747
  • Curated reference: UniProt P9WGU5 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 35 functional partner(s); context anchor oppB
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001370|Rv1280c|oppA
MADRGQRRGCAPGIASALRASFQGKSRPWTQTRYWAFALLTPLVVAMVLTGCSASGTQLELAPTADRRAAVGTTSDINQQDPATLQDGGNLRLSLTDFPPNFNILHIDGNNAEVAAMMKATLPRAFIIGPDGSTTVDTNYFTSIELTRTAPQVVTYTINPEAVWSDGTPITWRDIASQIHAISGADKAFEIASSSGAERVASVTRGVDDRQAVVTFAKPYAEWRGMFAGNGMLLPASMTATPEAFNKGQLDGPGPSAGPFVVSALDRTAQRIVLTRNPRWWGARPRLDSITYLVLDDAARLPALQNNTIDATGVGTLDQLTIAARTKGISIRRAPGPSWYHFTLNGAPGSILADKALRLAIAKGIDRYTIARVAQYGLTSDPVPLNNHVFVAGQDGYQDNSGVVAYNPEQAKRELDALGWRRSGAFREKDGRQLVIRDLFYDAQSTRQFAQIAQHTLAQIGVKLELQAKSGSGFFSDYVNVGAFDIAQFGWVGDAFPLSSLTQIYASDGESNFGKIGSPQIDAAIERTLAELDPGKARALANQVDELIWAEGFSLPLTQSPGTVAVRSTLANFGATGLADLDYTAIGFMRR