oppB Family assigned · medium auto-curated

H37Rv Rv1283c · MTBC0 mtbc0_001373 · 325 aa · 1445123–1446100 (-) · RefSeq NP_215799.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)oligopeptide ABC transporter permease OppB
MTBC0 PGAP re-annotationABC transporter permease
Revised (this work)ABC transporter permease. Pfam: BPD_transp_1_N (PF19300.5), BPD_transp_1 (PF00528.28).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WFZ7 SwissProt · reviewed · Evidence at protein level
UniProt nameOligopeptide transport system permease protein OppB
Curated functionPart of the ABC transporter complex OppABCD involved in the uptake of oligopeptides. Responsible for the translocation of the substrate across the membrane. Glutathione (GSH) uptake by mycobacteria through the OppABCD system contributes to the depletion of the GSH pool in infected macrophages, which impairs the ability of the macrophage to detoxify methylglyoxal (MG) and contributes to enhanced production of inflammatory cytokines. May also promote Mn(2+) uptake.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category E Amino acid transport and metabolism
P Inorganic ion transport and metabolism
Preferred nameoppB
eggNOG descriptiontransport
Orthologous groupCOG0601
KEGG orthology K02033
KEGG pathways map02024
KEGG modules M00239
Gene Ontology (14) GO:0003674, GO:0005215, GO:0005575, GO:0005623, GO:0005886, GO:0006810, GO:0008150, GO:0016020, GO:0022857, GO:0044464, GO:0051179, GO:0051234 +2 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.191 · strong purifying
Polymorphic sites (≥ 0.1% of strains) 4 synonymous, 2 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
BPD_transp_1_NPF19300.5 1.8e-051–76 Binding-prot-dependent transport system membrane comp, N-term
BPD_transp_1PF00528.28 3.1e-36113–323 Binding-protein-dependent transport system inner membrane component

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: oppC (oligopeptide ABC transporter permease OppC), high confidence from genomic context alone (score 999 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1282c oppC oligopeptide ABC transporter permease OppC 999 999 ctx neighborhood:882 cooccurence:774 coexpression:670 database:900 textmining:591
Rv1280c oppA oligopeptide ABC transporter substrate-binding lipoprotein OppA 999 998 ctx neighborhood:881 cooccurence:693 coexpression:459 database:900 textmining:914
Rv1281c oppD oligopeptide ABC transporter ATP-binding protein OppD 998 997 ctx neighborhood:816 cooccurence:773 coexpression:404 database:900
Rv3663c dppD dipeptide ABC transporter ATP-binding protein DppD 997 989 ctx cooccurence:765 coexpression:473 database:900 textmining:791
Rv2585c lipoprotein 967 956 coexpression:459 database:900
Rv3664c dppC dipeptide ABC transporter permease DppC 981 937 ctx cooccurence:774 coexpression:670 textmining:711
Rv3666c dppA dipeptide ABC transporter substrate-binding lipoprotein DppA 881 778 ctx cooccurence:550 coexpression:470 textmining:487
Rv1284 canA beta-carbonic anhydrase 756 756 ctx neighborhood:756
Rv1166 lpqW monoacyl phosphatidylinositol tetramannoside-binding protein LpqW 649 627 coexpression:457
Rv2326c ABC transporter ATP-binding protein 623 588 coexpression:472
Rv1286 cysC adenylyl-sulfate kinase 430 409
Rv3662c hyp hypothetical protein 582 151 textmining:528

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: oligopeptide ABC transporter permease OppB
  • MTBC0 PGAP product: ABC transporter permease
  • Pfam (hmmscan --cut_ga): BPD_transp_1_N PF19300.5 (E=2e-05), BPD_transp_1 PF00528.28 (E=3e-36)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215799.1)
  • Domains: Pfam-A via hmmscan --cut_ga — BPD_transp_1_N (PF19300.5), BPD_transp_1 (PF00528.28)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0601
  • Curated reference: UniProt P9WFZ7 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 12 functional partner(s); context anchor oppC
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001373|Rv1283c|oppB
MTRYLARRLLNYLVLLALASFLTYCLTSLAFSPLESLMQRSPRPPQAVIDAKAHDLGLDRPILARYANWVSHAVRGDFGTTITGQPVGTELGRRIGVSLRLLVVGSVFGTVAGVVIGAWGAIRQYRLSDRVMTTLALLVLSTPTFVVANLLILGALRVNWAVGIQLFDYTGETSPGVAGGVWDRLGDRLQHLILPSLTLALAAAAGFSRYQRNAMLDVLGQDFIRTARAKGLTRRRALLKHGLRTALIPMATLFAYGVAGLVTGAVFVEKIFGWHGMGEWMVRGISTQDTNIVAAITVFSGAVVLLAGLLSDVIYAALDPRVRVS