kdpA Family assigned · medium auto-curated

H37Rv Rv1029 · MTBC0 mtbc0_001108 · 571 aa · 1159349–1161064 (+) · RefSeq NP_215545.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)potassium-transporting ATPase subunit A
MTBC0 PGAP re-annotationpotassium-transporting ATPase subunit KdpA
Revised (this work)Potassium-transporting ATPase subunit KdpA. Pfam: KdpA (PF03814.22).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WKF3 SwissProt · reviewed · Evidence at protein level
UniProt namePotassium-transporting ATPase potassium-binding subunit
Curated functionPart of the high-affinity ATP-driven potassium transport (or Kdp) system, which catalyzes the hydrolysis of ATP coupled with the electrogenic transport of potassium into the cytoplasm. This subunit binds the extracellular potassium ions and delivers the ions to the membrane domain of KdpB through an intramembrane tunnel.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category P Inorganic ion transport and metabolism
Preferred namekdpA
eggNOG descriptionPart of the high-affinity ATP-driven potassium transport (or Kdp) system, which catalyzes the hydrolysis of ATP coupled with the electrogenic transport of potassium into the cytoplasm. This subunit binds and transports the potassium across the cytoplasmic membrane
Orthologous groupCOG2060
EC number EC 3.6.3.12
KEGG orthology K01546
KEGG pathways map02020
Gene Ontology (62) GO:0003674, GO:0003824, GO:0005215, GO:0005488, GO:0005575, GO:0005618, GO:0005623, GO:0005886, GO:0006810, GO:0006811, GO:0006812, GO:0006813 +50 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.446 · purifying
Polymorphic sites (≥ 0.1% of strains) 7 synonymous, 9 missense, 0 nonsense, 2 frameshift
Disruption 2 distinct premature-stop/frameshift site(s); most common in 0.24% of strains (349) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
KdpAPF03814.22 8.0e-24011–568 Potassium-transporting ATPase A subunit

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: kdpC (potassium-transporting ATPase subunit C), high confidence from genomic context alone (score 1000 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv1031 kdpC potassium-transporting ATPase subunit C 999 1000 ctx neighborhood:881 fusion:817 cooccurence:774 coexpression:843 experimental:895 database:900 textmining:833
Rv1030 kdpB potassium-transporting ATPase subunit B 999 1000 ctx neighborhood:881 cooccurence:774 coexpression:976 experimental:895 database:900 textmining:926
Rv1028c kdpD sensor protein KdpD 989 978 ctx neighborhood:747 cooccurence:772 coexpression:653 textmining:522
Rv1028A kdpF membrane protein KdpF 944 883 ctx neighborhood:882 textmining:541
Rv1027c kdpE transcriptional regulator KdpE 927 876 ctx neighborhood:747 textmining:437
Rv1033c trcR two component transcriptional regulator TrcR 423 391
Rv0410c pknG serine/threonine-protein kinase PknG 680 63 textmining:673
Rv1469 ctpD cobalt/nickel-exporting P-type ATPase 494 52 textmining:488
Rv0425c ctpH metal cation transporting ATPase H 521 51 textmining:517
Rv0103c ctpB cation-transporter P-type ATPase B 466 50 textmining:461
Rv2109c prcA proteasome subunit alpha 430 44 textmining:429
Rv0812 4-amino-4-deoxychorismate lyase 701 41 textmining:701
Rv0561c menJ oxidoreductase 583 41 textmining:584
Rv0107c ctpI cation-transporter ATPase I 515 41 textmining:516
Rv2110c prcB proteasome subunit beta 434 41 textmining:435

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: potassium-transporting ATPase subunit A
  • MTBC0 PGAP product: potassium-transporting ATPase subunit KdpA
  • Pfam (hmmscan --cut_ga): KdpA PF03814.22 (E=8e-240)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215545.1)
  • Domains: Pfam-A via hmmscan --cut_ga — KdpA (PF03814.22)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2060
  • Curated reference: UniProt P9WKF3 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 15 functional partner(s); context anchor kdpC
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001108|Rv1029|kdpA
MSGTSWLQFAALIAVLLLTAPALGGYLAKIYGDEAKKPGDRVFGPIERVIYQVCRVDPGSEQRWSTYALSVLAFSVMSFLLLYGIARFQGVLPFNPTDKPAVTDHVAFNAAVSFMTNTNWQSYSGEATMSHFTQMTGLAVQNFVSASAGMCVLAALIRGLARKRASTLGNFWVDLARTVLRIMFPLSFVVAILLVSQGVIQNLHGFIVANTLEGAPQLIPGGPVASQVAIKQLGTNGGGFFNVNSAHPFENYTPIGNFVENWAILIIPFALCFAFGKMVHDRRQGWAVLAIMGIIWIGMSVAAMSFEAKGNPRLDALGVTQQTTVDQSGGNLEGKEVRFGVGASGLWAASTTGTSNGSVNSMHDSYTPLGGMVPLAHMMLGEVSPGGTGVGLNGLLVMAILAVFIAGLMVGRTPEYLGKKIQATEMKLVTLYILAMPIALLSFAAASVLISSALASRNNPGPHGLSEILYAYTSGANNNGSAFAGLTASTWSYDTTIGVAMLIGRFFLIIPVLAIAGSLARKGTTPVTAATFPTHKPLFVGLVIGVVLIVGGLTFFPALALGPIVEQLSTQ