Rv1024 Family assigned · medium auto-curated

H37Rv Rv1024 · MTBC0 mtbc0_001100 · 228 aa · 1153195–1153881 (+) · RefSeq NP_215540.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)membrane protein
MTBC0 PGAP re-annotationseptum formation initiator family protein
Revised (this work)Septum formation initiator family protein. Pfam: DivIC (PF04977.21).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P96376 TrEMBL · unreviewed · Predicted
UniProt namePossible conserved membrane protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category D Cell cycle control, cell division, chromosome partitioning
Preferred namedivIC
eggNOG descriptionSeptum formation initiator
Orthologous groupCOG2919
KEGG orthology K13052

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.273 · purifying
Polymorphic sites (≥ 0.1% of strains) 4 synonymous, 3 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
DivICPF04977.21 4.5e-1986–161 Septum formation initiator

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: eno (enolase), high confidence from genomic context alone (score 904 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv2151c ftsQ cell division protein FtsQ 983 955 experimental:928 textmining:655
Rv1023 eno enolase 903 904 ctx neighborhood:882
Rv1025 hyp hypothetical protein 885 885 ctx neighborhood:861
Rv1026 ppx2 hyp hypothetical protein 909 871 ctx neighborhood:861
Rv1020 mfd transcription-repair coupling factor 776 776 ctx neighborhood:505 cooccurence:545
Rv1022 lpqU lipoprotein LpqU 775 776 ctx neighborhood:774
Rv0007 membrane protein 771 771 ctx cooccurence:768
Rv3909 hyp hypothetical protein 768 768 ctx cooccurence:765
Rv3835 hyp hypothetical protein 767 768 ctx cooccurence:761
Rv3658c transmembrane protein 747 748 ctx cooccurence:746
Rv0538 membrane protein 744 745 ctx cooccurence:741
Rv2843 hyp hypothetical protein 729 729 ctx cooccurence:725
Rv2164c hyp hypothetical protein 708 696 ctx cooccurence:688
Rv3258c hyp hypothetical protein 668 668 ctx cooccurence:666
Rv1648 transmembrane protein 665 665 ctx cooccurence:664

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: membrane protein
  • MTBC0 PGAP product: septum formation initiator family protein
  • Pfam (hmmscan --cut_ga): DivIC PF04977.21 (E=5e-19)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215540.1)
  • Domains: Pfam-A via hmmscan --cut_ga — DivIC (PF04977.21)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG2919
  • Curated reference: UniProt P96376 (TrEMBL, unreviewed; Predicted)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 53 functional partner(s); context anchor eno
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001100|Rv1024|
MPEAKRPESKRRSPASRPGKAGDSVRGGRATKPSAKPSTPAPHASRKTTRTPHEHIVEPIKRAITESVEKRSEQRLGFTARRAAILAAVVCVLTLTIARPVRTYFAQRAEMEQLAATEAMLRRQIADLEEQQVKLADPAYIAAQARERLGFVMPGDIPFQVQLPSTPLAPPQPGSDAATATNNEPWYTALWHTIADDPHLPPAAPPAPEPGRPGPLPPASPNPEQPGG