pgi Resolved · high auto-curated

H37Rv Rv0946c · MTBC0 mtbc0_001009 · 553 aa · 1062239–1063900 (-) · RefSeq NP_215461.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)glucose-6-phosphate isomerase
MTBC0 PGAP re-annotationglucose-6-phosphate isomerase
Revised (this work)Glucose-6-phosphate isomerase. Pfam: PGI (PF00342.25).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P9WN69 SwissProt · reviewed · Evidence at protein level
UniProt nameGlucose-6-phosphate isomerase
EC (curated) EC 5.3.1.9
Curated functionCatalyzes the reversible isomerization of glucose-6-phosphate to fructose-6-phosphate.

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category G Carbohydrate transport and metabolism
Preferred namepgi
eggNOG descriptionBelongs to the GPI family
Orthologous groupCOG0166
EC number EC 5.3.1.9
KEGG orthology K01810
KEGG pathways map00010, map00030, map00500, map00520, map01100, map01110, map01120, map01130, map01200
KEGG modules M00001, M00004, M00114
Gene Ontology (122) GO:0003674, GO:0003824, GO:0004347, GO:0005575, GO:0005623, GO:0005886, GO:0005975, GO:0006082, GO:0006090, GO:0006091, GO:0006096, GO:0006139 +110 more

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains)

pN/pS 0.34 · purifying
Polymorphic sites (≥ 0.1% of strains) 5 synonymous, 5 missense, 0 nonsense, 0 frameshift

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
PGIPF00342.25 9.3e-20054–542 Phosphoglucose isomerase

Functional interaction network (STRING v12, guilt-by-association)

PartnerProductScoreNo text-miningChannels (≥400)
Rv3010c pfkA 6-phosphofructokinase 995 981 coexpression:779 database:900 textmining:748
Rv1438 tpi triosephosphate isomerase 995 979 coexpression:851 database:800 textmining:804
Rv3068c pgmA phosphoglucomutase PgmA 990 979 coexpression:749 database:900 textmining:590
Rv1449c tkt transketolase 988 974 coexpression:675 database:900 textmining:564
Rv1617 pykA pyruvate kinase 991 970 coexpression:822 database:800 textmining:719
Rv1448c tal transaldolase 991 968 coexpression:642 database:900 textmining:755
Rv1023 eno enolase 993 966 coexpression:799 database:800 textmining:819
Rv1436 gap glyceraldehyde 3-phosphate dehydrogenase 981 955 coexpression:724 database:800 textmining:620
Rv1447c zwf2 glucose-6-phosphate 1-dehydrogenase 977 952 coexpression:480 database:900 textmining:550
Rv1121 zwf1 glucose-6-phosphate 1-dehydrogenase 974 952 coexpression:482 database:900 textmining:497
Rv0363c fba fructose-bisphosphate aldolase 978 949 coexpression:735 database:800 textmining:596
Rv3255c manA mannose-6-phosphate isomerase 944 927 database:900
Rv0489 gpm1 2,3-bisphosphoglycerate-dependent phosphoglycerate mutase 959 920 coexpression:614 database:800 textmining:511
Rv2029c pfkB 6-phosphofructokinase PfkB 954 909 database:900 textmining:518
Rv3436c glmS glucosamine--fructose-6-phosphate aminotransferase 958 908 database:900 textmining:562

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: glucose-6-phosphate isomerase
  • MTBC0 PGAP product: glucose-6-phosphate isomerase
  • Pfam (hmmscan --cut_ga): PGI PF00342.25 (E=9e-200)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_215461.1)
  • Domains: Pfam-A via hmmscan --cut_ga — PGI (PF00342.25)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0166
  • Curated reference: UniProt P9WN69 (SwissProt, reviewed; Evidence at protein level)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 120 functional partner(s)
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_001009|Rv0946c|pgi
MTSAPIPDITATPAWDALRRHHDQIGNTHLRQFFADDPGRGRELTVSVGDLYIDYSKHRVTRETLALLIDLARTAHLEERRDQMFAGVHINTSEDRAVLHTALRLPRDAELVVDGQDVVTDVHAVLDAMGAFTDRLRSGEWTGATGKRISTVVNIGIGGSDLGPVMVYQALRHYADAGISARFVSNVDPADLIATLADLDPATTLFIVASKTFSTLETLTNATAARRWLTDALGDAAVSRHFVAVSTNKRLVDDFGINTDNMFGFWDWVGGRYSVDSAIGLSLMTVIGRDAFADFLAGFHIIDRHFATAPLESNAPVLLGLIGLWYSNFFGAQSRTVLPYSNDLSRFPAYLQQLTMESNGKSTRADGSPVSADTGEIFWGEPGTNGQHAFYQLLHQGTRLVPADFIGFAQPLDDLPTAEGTGSMHDLLMSNFFAQTQVLAFGKTAEEIAADGTPAHVVAHKVMPGNRPSTSILASRLTPSVLGQLIALYEHQVFTEGVVWGIDSFDQWGVELGKTQAKALLPVITGAGSPPPQSDSSTDGLVRRYRTERGRAG