Rv0024 Family assigned · medium auto-curated

H37Rv Rv0024 · MTBC0 mtbc0_000029 · 281 aa · 28344–29189 (+) · RefSeq NP_214538.1

Annotation: from legacy to revised

Legacy (H37Rv / Mycobrowser)NLP/P60 family protein
MTBC0 PGAP re-annotationC40 family peptidase
Revised (this work)C40 family peptidase. Pfam: NLPC_P60 (PF00877.26).

Auto-curated: this verdict and function were generated by rules from PGAP + Pfam + Foldseek and have not been hand-reviewed.

Curated reference (UniProt)

UniProt P71594 TrEMBL · unreviewed · Inferred from homology
UniProt nameSecreted protein P60-related protein

Functional vocabulary (eggNOG-mapper, orthology transfer)

COG category M Cell wall / membrane / envelope biogenesis
eggNOG descriptionNlpC/P60 family
Orthologous groupCOG0791
KEGG orthology K21471

Orthology-based transfer (eggNOG 5.0.2, diamond). EC/KO/GO/CAZy are computed annotations, not manual curation; cross-check against the primary literature before treating a specific reaction as established.

Conservation & selection (intra-MTBC, 145 209 strains) pseudogene candidate

pN/pS 0.268 · purifying
Polymorphic sites (≥ 0.1% of strains) 6 synonymous, 4 missense, 0 nonsense, 2 frameshift
Disruption 2 distinct premature-stop/frameshift site(s); most common in 26.25% of strains (38111) · clonal

pN/pS from segregating SNPs (singletons removed) normalised by possible sites. Low pN/pS = purifying selection (a strong signal that a "hypothetical" is a real, constrained gene). A high pN/pS is ambiguous: relaxed constraint or positive selection (drug resistance, antigenic variation) inflate it; e.g. rpoB/katG/pncA score high here for resistance, not loss of function. A clonal disruption (one allele over a clade) suggests lineage pseudogenisation; a convergent one (many independent alleles) is typical of resistance loss-of-function.

Domains (Pfam, hmmscan --cut_ga)

PfamAccessioni-EvalueResiduesDescription
NLPC_P60PF00877.26 7.0e-22182–272 NlpC/P60 family

Functional interaction network (STRING v12, guilt-by-association)

Closest characterised functional partner: Rv0023 (transcriptional regulator), high confidence from genomic context alone (score 826 excluding text-mining).

PartnerProductScoreNo text-miningChannels (≥400)
Rv0023 transcriptional regulator 825 826 ctx neighborhood:816
Rv0025 hyp hypothetical protein 727 728 ctx neighborhood:716
Rv0022c whiB5 transcriptional regulator WhiB5 630 617 ctx neighborhood:615
Rv0026 hyp hypothetical protein 446 446 ctx neighborhood:438
Rv1477 ripA peptidoglycan endopeptidase RipA 511 358
Rv2190c ripC endopeptidase 465 308
Rv3433c nnr bifunctional ADP-dependent (S)-NAD(P)H-hydrate dehydratase/NAD(P)H-hydrate epimerase 407 284
Rv2974c hyp hypothetical protein 419 188
Rv3915 cwlM peptidoglycan hydrolase 674 147 textmining:634
Rv2704 hyp hypothetical protein 463 100 textmining:429
Rv2389c rpfD resuscitation-promoting factor RpfD 549 86 textmining:528
Rv1337 integral membrane protein 664 77 textmining:651
Rv1009 rpfB resuscitation-promoting factor RpfB 582 71 textmining:569
Rv2864c penicillin-binding lipoprotein 817 66 textmining:812
Rv1430 PE16 PE family protein PE16 534 63 textmining:523

STRING combines evidence channels (neighborhood, fusion, cooccurrence, coexpression, experimental, database, text-mining) into a 0–1000 score. The ctx badge marks edges carried by the genomic-context channels (conserved neighborhood, fusion, phylogenetic co-occurrence), which are independent of orthology and structure and the strongest signal for an unknown gene. The no text-mining column recomputes the score from data alone, so a link that does not depend on the literature is visible. Association is a function hypothesis, not proof: corroborate with the operon context and the primary literature before assigning a function.

Evidence

  • Legacy H37Rv annotation: NLP/P60 family protein
  • MTBC0 PGAP product: C40 family peptidase
  • Pfam (hmmscan --cut_ga): NLPC_P60 PF00877.26 (E=7e-22)
  • (auto-curated by rules from PGAP + Pfam + Foldseek; not hand-reviewed)

Sources

  • Ancestral sequence & coordinates: Harrison LB et al. (2024), An imputed ancestral reference genome for the MTBC, doi:10.1101/2023.09.07.556366
  • Product annotation: NCBI PGAP on MTBC0; legacy from H37Rv NC_000962.3 (RefSeq NP_214538.1)
  • Domains: Pfam-A via hmmscan --cut_ga — NLPC_P60 (PF00877.26)
  • Sequence-level signal: ESM Atlas (EvolutionaryScale × BioHub) — exploratory
  • Controlled vocabulary: eggNOG-mapper 2.1.12 (Cantalapiedra et al. 2021, doi:10.1093/molbev/msab293), eggNOG 5.0 DB (Huerta-Cepas et al. 2019) — OG COG0791
  • Curated reference: UniProt P71594 (TrEMBL, unreviewed; Inferred from homology)
  • Intra-MTBC selection: pN/pS and disruption from SPDI variants of 145 209 MTBC strains (this work, local collection vs H37Rv NC_000962.3)
  • Interaction network: STRING v12.0 (Szklarczyk et al. 2023, doi:10.1093/nar/gkac1000), taxon 83332, CC-BY 4.0 — 20 functional partner(s); context anchor Rv0023
  • Primary literature: none located yet; annotation rests on the domain/homology sources above.

Ancestral MTBC0 protein sequence

>mtbc0_000029|Rv0024|
MNYSEVELLSRAHQLFAGDSRRPGLDAGTTPYGDLLSRAADLNVGAGQRRYQLAVDHSRAALLSAARTDAAAGAVITGAQRDRAWARRSTGTVLDEARSDTTVTAVMPIAQREAIRRRVARLRAQRAHVLTARRRARRHLAALRALRYRVAHGPGVALAKLRLPSPSGRAGIAVHAALSRLGRPYVWGATGPNQFDCSGLVQWAYAQAGVHLDRTTYQQINEGIPVPRSQVRPGDLVFPHPGHVQLAIGNNLVVEAPHAGASVRVSSLGNNVQIRRPLSGR